There is so great a difference between the nervous system of the
naked and of the covered eyed Medusæ, that a simultaneous description
of the nervous system in both groups is not by these authors
considered practicable. Beginning, therefore, with the naked-eyed
division, they describe the nervous system as here consisting of two
parts, a central and a peripheral. The central part is localized in
the margin of the swimming-bell, and there forms a "nerve-ring,"
which is divided by the insertion of the "veil"[3] into an upper and
a lower nerve-ring. In many species the upper nerve-ring is spread
out in the form of a flattish layer, which is somewhat thickened
where it is in contact with the veil. In these species the nerve-ring
is only indistinctly marked off from the surrounding tissues. But
in other species the crowding together of the nerve-fibres at the
insertion of the veil gives rise to a considerable concentration
of nervous structures; while in others, again, this concentration
proceeds to the extent of causing a well-defined swelling of nervous
tissue against the epithelium of the veil and umbrella. In the
Geryonidæ this swelling is still further strengthened by a peculiar
modification of the other tissues in the neighbourhood, which had
been previously described by Professor Haeckel. In all species the
upper nerve-ring lies entirely in the ectoderm. Its principal mass
is composed of nerve-fibres of wonderful tenuity, among which are
to be found sparsely scattered ganglion-cells. The latter are for
the most part bi-polar, more seldom multi-polar. The fibres which
emanate from them are very delicate, and, becoming mixed with others,
do not admit of being further traced. Where the nervous tissue meets
the enveloping epithelium it is connected with the latter from
within, but differs widely from it; for the nerve-cells contain a
longitudinally striated cylindrical or thread-like nucleus which
carries on its peripheral end a delicate hair, while its central end
is prolonged into a fine nerve-fibre. There are, besides these, two
other kinds of cells which form a transition between the ganglion and
the epithelium cells. The first kind are of a long and cylindrical
form, the free ends of which reach as far as the upper surface of the
epithelium The second kind lie for the most part under the upper
surface. They are of a large size, and present, coursing towards
the upper surface, a long continuation, which at its free extremity
supports a hair. In some cases this continuation is smaller, and
stops short before reaching the outer surface. Drs. Hertwig observe
that in these peculiar cells we have tissue elements which become
more and more like the ordinary ganglion-cells of the nerve-ring the
more that their long continuation towards the surface epithelium
is shortened or lost, and these authors are thus led to conclude
that the upper nerve-ring was originally constituted only by such
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