I may give one other experiment having the same object and tendency.
Employing single induction shocks of slightly more than minimal
intensity, and throwing them in at twice the rate that was required
to produce a strong response to every shock, I found that midway
between every two strong responses there was a weak response. In
other words, a stimulus of uniform intensity gives rise alternately
to a strong and to a weak contraction, as shown in the appended
tracing (Fig. 27). It will be observed that in this tracing each
large curve represents the whole time occupied by the strong
contraction, the latter beginning at the highest point of the
curve on the left-hand side in each case. The effect of the weak
contraction is that of momentarily interrupting the even sweep of
diastole after the strong contraction, and therefore the result on
the tracing is a slight depression in the otherwise even curve of
ascent. Lest any doubt should arise from the smallness of the curves
representing the weak contractions that the former are in some
way accidental, I may draw attention to the fact that the period
of latent stimulation is the same in the case of all the curves.
To render this apparent, I have placed crosses below the smaller
curves, which show in each case the exact point where the depressing
effect of these smaller curves on the ascending sweeps of the larger
curves first become apparent--_i.e._ the point at which the feeble
contraction begins. Now, what I wish to be gathered from the whole
tracing is this. If the strength of the induction shocks had been
much greater than it was, _all_ the contractions would have become
strong contractions, and tetanus would have been the result. But,
as the strength of the induction shocks was only slightly more than
minimal, the exhaustion consequent on every strong contraction so
far diminished the irritability of the tissue that when, during
the process of relaxation, another shock _of the same intensity_
was thrown in, the stimulus was only strong enough, in relation
to the diminished irritability of the partly recovered tissue, to
cause a feeble contraction. And these facts tend still further to
substantiate the hypothesis whereby I have sought to explain the
phenomena of artificial rhythm.
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