[4] See "Observations on the Nervous System of _Aurelia aurita_,"
_Phil. Trans._, pt. ii., 1878.
Lastly, returning to the researches of Drs. Hertwig, these authors
compare the nervous system of the naked-eyed with that of the
covered-eyed Medusæ, with the view of indicating the points which
show the latter to be less developed than the former. These points
are, that in the nerve-centres of the covered-eyed Medusæ there
are no true ganglion-cells, or only very few; that the mass of the
central nervous system is very small; and that the centralization
of the nervous system is less complete in the one group than in the
other. In their memoir these authors further supply much interesting
information touching the structure of the sense-organs in various
species of Medusæ; but it seems scarcely necessary to extend the
present _résumé_ of their work by entering into this division of
their subject.
In a later publication, entitled "Der Organismus der Medusen und
seine Stellung zur Keimblättertheorie," Drs. Hertwig treat of
sundry features in the morphology of the Medusæ which are of great
theoretical importance; but here again it would unduly extend the
limits of the present treatise if I were to include all the ground
which has been so ably cultivated by these industrious workers.
It will presently be seen in how striking a manner all the
microscopical observations to which I have now briefly alluded are
confirmed by the physiological observations--or, more correctly, I
might say that the microscopical observations, in so far as they
were concerned with demonstrating the existence of nerve-tissue in
the Medusæ, were forestalled by these physiological experiments;
for, with the exception of Professor Haeckel's work on Geryonidæ,
they were all of later publication. But in matters of scientific
inquiry mere priority is not of so much importance as it is too often
supposed to be. Thus, in the present instance, no one of the workers
was in any way assisted by the publications of another. In each case
the work was independent and almost simultaneous.
The remark just made applies also to the only research which still
remains to be mentioned. This is the investigation undertaken and
published by Professor Eimer.[5] He began, like myself, by what in
the next chapter I call the "fundamental observation" on the effects
of excising the nerve-centres, and from this basis he worked both at
the physiology and the morphology of the neuro-muscular tissues. In
point of time, I was the first to make the fundamental observation,
and he was the first to publish it. The sundry features in which our
subsequent investigations agreed, and those in which they differed, I
shall mention throughout the course of the following pages.
[5] "Die Medusen physiologisch und morphologisch auf ihr
Nervensystem untersucht" (Tübingen, 1878).
* * * * *
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