very little more stable than it is, it would altogether fail to be
thrown down by the luminous vibrations, or would occupy so long a
time in the process that the visual sense would be of no use to its
possessor. How great is the contrast between the excitability of
such a sense-organ and that of a fully evolved eye, which is able
to effect the needful molecular changes in response to a flash as
instantaneous as that of lightning.
[10] The period of latent stimulation merely means the time
after the occurrence of an excitation during which a series of
physiological processes are taking place, which terminate in
a contraction; so that, whether the excitation is of a strong
or of a weak intensity, the period of latent stimulation is
not much affected. The above question, therefore, was simply
this--Does the prolonged delay on the part of these ganglia, in
responding to light, represent the time during which the series
of physiological processes are taking place in response to an
adequate stimulus, or does it represent the time during which
light requires to act before it becomes an adequate stimulus?
With regard to luminous stimulation, it is only necessary further to
observe that responses were given equally well to direct sunlight,
diffused daylight, and to light reflected from a mirror inclined at
the polarizing angle. It must also be stated that responses are given
to any of the luminous rays of the spectrum when these are employed
separately; but that neither the non-luminous rays beyond the red,
nor those beyond the violet, appear to exert the smallest degree of
stimulating influence.
_Electrical Stimulation._
All the excitable parts of all the Medusæ which I have examined are
highly sensitive to electrical stimulation, both of the constant and
of the induced current.
Exploration with needle-point terminals and induction shocks of
graduated strength showed that certain parts or tracts of the
nectocalyx are more sensitive than others. The most sensitive
parts are those which correspond with the distribution of the main
nerve-trunks, _i.e._ round the margin of the nectocalyx and along
the course of the radial tubes. The external or convex surface of a
nectocalyx or umbrella is totally insensitive to stimulation, and
the same statement applies to the whole thickness of the gelatinous
substance to which the neuro-muscular sheet is attached.
Public-domain text, read in full here on John Shaqi.
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