acquainted with the method, when it is stated that the upper line is
a record of the movements of the electro-magnet writer, and the lower
line that of the movements of the other writer. It will be observed
that the point _a_ in the upper line marks the point at which the
induction shock was thrown in; so that by first producing the
perpendicular till it meets the lower line at _b_, and then measuring
the distance between the point _b_ and the point _c_, at which the
curve in the lower line first begins, the latent period (_b c_) is
determined--the time occupied by the rotation of the cylinder from
_b_ to _c_ being known.
[12] It may here be stated that in all the experiments on
stimulation subsequently to be detailed, there is no difference
to be observed between the behaviour of an entire swimming organ
deprived of its ganglia, and that of a portion of any size which
may be separated from it.
_Summation of Stimuli._--In this way I have been able to ascertain
the period of latent stimulation in Aurelia aurita with accuracy.
It must be stated at the outset, however, that the period is subject
to great variations under certain varying conditions, so that we can
only arrive at a just estimation of it by understanding the nature
of the modifying causes. To take the simplest cause first, suppose
that the paralyzed Aurelia has been left quiet for several minutes in
sea-water at forty-five degrees, and that it is then stimulated by
means of a single induction shock; the responsive contraction will
be comparatively feeble with a very long period of latency, viz.
five-eighths of a second. If another shock of the same intensity be
thrown in as soon as the tissue has relaxed, a somewhat stronger
contraction, with a somewhat shorter latent period, will be given.
If the process is again repeated, the response will be still more
powerful, with a still shorter period of latency; and so on, perhaps,
for eight or ten stages, when the maximum force of contraction of
which the tissue is capable will have been attained, while the period
of latency will have been reduced to its minimum. This period is
three-eighths of a second, or, in some cases, slightly less.
Now, we have here a remarkable series of phenomena, and as it is
a series which never fails to occur under the conditions named,
I append tracings to give a better idea of the very marked and
striking character of the results. The first tracing (Fig. 2) is a
record of the successive increments of the responses to successive
induction shocks of the same intensity, thrown in at three seconds'
intervals--the cylinder being stationary during each response, and
rotated a short distance with the hand during each interval of repose.
[Illustration: Fig. 2.]
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