Lastly, the third mode of section is represented in the next cut.
Here seven of the marginal ganglia having been removed as before,
the eighth one was made the point of origin of a circumferential
section, which was then carried round and round the bell in the
form of a continuous spiral--the result, of course, being this long
ribbon-shaped strip of tissue with the ganglion at one end and the
remainder of the swimming-bell at the other. Well, as before, the
contraction-waves always originated at the ganglion; but now they
had to course all the way along the strip until they arrived at its
other extremity; and, as each wave arrived at that extremity, it
delivered its influence into the remainder of the swimming-bell,
which thereupon contracted. Now, in this experiment, when the spiral
strip is only made about half an inch broad, it may be made more than
a yard long before all the bell is used up in making the strip; and
as nothing can well be imagined as more destructive of the continuity
of a nerve-plexus than this spiral mode of section must be, we cannot
but regard it as a very remarkable fact that the nerve-plexus should
still continue to discharge its function. Indeed, so remarkable
does this fact appear, that to avoid accepting it we may well
feel inclined to resort to another hypothesis, namely, that these
contraction-waves do not depend for their passage on the nervous
network at all, but that they are of the nature of the muscle-waves,
or of the waves which we see in undifferentiated protoplasm,
where all parts of the mass being equally excitable and equally
contractile, however severely we cut the mass, as long as we do not
actually divide it, contraction-waves will pass throughout the whole
mass. But this very reasonable hypothesis of the contraction-waves in
the Medusæ being possibly nothing more than muscle-waves is negatived
by other facts, which I shall now proceed to state.
Public-domain text, read in full here on John Shaqi.
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