muscle-waves, so long as the _conduction_ or _undoubtedly nervous_
waves are equally able to pass round sections interposed in their
path. Briefly, then, I find that the nervous waves of stimulation
are quite as able to pass round these interposed sections as are the
waves of contraction. Thus, for instance, in this specimen (Fig. 12),
the tentacular wave of stimulation continued to pass as before, even
after I had submitted the parallelogram of tissue to the tremendously
severe form of section which is represented in the illustration;
and this fact, in my opinion, is one of the most important that has
been brought to light in the whole range of invertebrate physiology.
For what does it prove? It proves that the distinguishing function
of nerve, where it first appears upon the scene of life, admits of
being performed vicariously to almost any extent by all parts of
the same tissue-mass. If we revert to our old illustration of the
muslin as representing the nerve-plexus, it is clear that, however
much we choose to cut the sheet of muslin with such radial or spiral
sections as are represented in the illustrations, one could always
trace the threads of the muslin with a needle round and round the
disc, without once interrupting the continuity of the tracing; for
on coming to the end of a divided thread, one could always double
back on it and choose another thread which might be running in the
required direction. And this is what we are now compelled to believe
takes place in the fibres of this nervous network, if we assume that
these visible fibres are the only conductile elements which are
present. Whenever a stimulus wave reaches a cut, we must conclude
that it doubles back and passes into the neighbouring fibres, and so
on, time after time, till it succeeds in passing round and round any
number of overlapping cuts.
Public-domain text, read in full here on John Shaqi.
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