In various modes of section of Aurelia I have several times observed
a fact that is worth recording. It sometimes happens that when the
connecting isthmus between two almost severed areas of excitable
tissue is very narrow, the passage of contraction-waves across the
isthmus depends upon the freshness, or freedom from exhaustion,
of the tissue which constitutes the isthmus. That is to say, on
faradizing one of the two tissue-areas which the isthmus serves
to connect, the resulting contraction-waves will at first pass
freely across the isthmus; but after a time it may happen in some
preparations that every now and then a contraction-wave fails to
pass across the isthmus. When this is the case, if the stimulation
is still continued, a greater and greater proportion of waves fail
to pass across the isthmus, until perhaps only one in every five or
six becomes propagated from the one area to the other. If single
induction-shocks be then substituted for the faradaic stimulation,
it may be found that by leaving an interval of four or five seconds
between the successive shocks, every wave that is started in the
one area will be propagated across the isthmus to the other area.
But if the interval between the successive shocks be reduced to
two or three seconds, every now and then a wave will fail to pass
across the isthmus; and if the interval be still further reduced to
one second, or half a second, comparatively few of the waves will
pass across. Now, however, if the tissue be allowed five minutes'
rest from stimulation, and the single shocks be thrown in at one
second's intervals, all the first six or ten waves will pass across
the isthmus, after which they begin to become blocked as before. It
may be observed also that when the waves are thus blocked, owing to
exhaustion of the connecting isthmus, they may again be made to force
a passage by increasing the intensity of the stimulation, and so
giving rise to stronger waves having a greater power of penetration.
Thus, on re-enforcing the electrical stimulus with the simultaneous
application of a drop of spirit, the resulting waves of contraction
are almost sure to pass across the isthmus, even though this has been
exhausted in the manner just described.
_Ganglia appearing to assert their Influence at a Distance from their
own Seat._
Another fact, which I have several times noticed during my sections
of Aurelia, also deserves to be recorded. I have observed it under
several modes of section, but it will be only necessary to describe
one particular case.
Public-domain text, read in full here on John Shaqi.
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