Journal of the Proceedings of the Linnean Society - Vol. 4: ZoologyLinnean Society of London
Science
Journal of the Proceedings of the Linnean Society - Vol. 4: Zoology
Linnean Society of London
Zoology -- Periodicals
There are twenty-seven or twenty-eight teeth in the upper, and
twenty-five or twenty-six in the lower jaw. The mandibular symphysis
extends to the twenty-third or twenty-fourth tooth. The lateral teeth
of both jaws are, all but the very hindmost, directed obliquely
downwards (or upwards), forwards or outwards, and are not received
into interdental pits. The anterior margins of the orbits are raised.
The premaxillæ and the end of the mandible are greatly expanded. The
premaxillo-maxillary suture reaches the level of the fourth tooth behind
the canine notch.
The only true _Gavialis_ is the well-known _G. Gangeticus_ from the
East Indies. In this 'Gavial,' or 'Garrhial,' the vomers are slender
bones which do not extend further forwards than the level of the
twenty-second or twenty-first tooth, and have but a very short and
slender representative of the anterior flattened division of the bone in
_Jacare_; posteriorly they extend back to the level of the descending
processes of the prefrontals. In a skull 25 inches long the vomers
have a length of about 4 inches, extending as they do a little further
forward than the palato-maxillary suture. The median nares are opposite
the twenty-fifth tooth.
* * * * *
All the _Crocodilia_ which I have enumerated are provided with two
perfectly distinct kinds of dermal armour,--the one consisting of
plates of horn, produced by a modification of the superficial layer of
the epidermis; the other composed of discs of bone marked by a peculiar
pitted sculpture on their outer surfaces, and developed within the
substance of the dermis. To the former I shall apply the term "scales;"
the latter are what I have denominated "scutes."
All recent _Crocodilia_ have both scales and scutes in the dorsal
region of the body, the scutes underlying, and having the same general
form as the scales. In all, the ventral region of the body is also
covered with scales which have a very definite shape; but in no recent
Crocodilian which I have examined, save those species which are
included in the genera _Caiman_ and _Jacare_, are there any scutes in
the ventral region.
Again, in the genera _Alligator_, _Crocodilus_, _Mecistops_,
Rhynchosuchus, and _Gavialis_, the edges of the scutes, except those of
the two median longitudinal rows, are hardly ever united by sutures,
nor do the posterior margins of those in each transverse row overlap the
anterior margins of the succeeding row. At any rate, there is no flat,
bevelled, articular facet on the outer surface of the anterior margin of
a scute, for articulation with the inner surface of the posterior margin
of its predecessor. In the genera _Caiman_ and _Jacare_, however, the
lateral edges of all the scutes of the dorsal and ventral shields are
united by serrated sutures; and the anterior end of the outer face of
each is provided with a well-marked smooth facet, which is overlapped by
the smooth under-surface of the scute in front of it.
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