It is a matter of regret that Greville should have so entirely ignored
lichens in his great work on _Scottish Cryptogams_[99]. The two species
of Lichina are the only ones he figured, and these he took to be algae.
He[100] was well acquainted with lichens, for in the _Flora Edinensis_ he
lists 128 species for the Edinburgh district, arranging the genera under
“Lichenes” with the exception of _Opegrapha_ and _Verrucaria_ which are
placed with the fungus genus _Poronia_ in “Hypoxyla.” Though he cites
the publications of Acharius, he does not employ his scientific terms,
possibly because he was writing his diagnoses in English. Two other
British works of this time still remain to be chronicled: Hooker’s[101]
contributions to Smith’s _English Flora_ and Taylor’s[102] work on
lichens in Mackay’s _Flora Hibernica_. Through these the knowledge of the
subject was very largely extended in our country.
The classification of lichens and their place in the vegetable kingdom
were now firmly established on the lines laid down by Acharius.
Fries[103] in his important work _Lichenographia Europaea_ more or
less followed his distinguished countryman. The uncertainty as to the
position and relationship of lichens had rendered the task of systematic
arrangement one of peculiar difficulty and had unduly absorbed attention;
but now that a satisfactory order had been established in the chaos
of forms, the way was clear for other aspects of the study. Several
writers expressed their views by suggesting somewhat different methods
of classification, others wrote monographs of separate groups, or
genera. Fée[104] published an Essay on the Cryptogams (mostly lichens)
that grew on officinal exotic barks; Flörke[105] took up the difficult
genus _Cladonia_; Wallroth[106] also wrote on _Cladonia_; Delise[107] on
_Sticta_, and Chevalier[108] published a long and elaborate account of
Graphideae.
Wallroth and Meyer at this time published, simultaneously, important
studies on the general morphology and physiology of lichens.
Wallroth[109] had contemplated an even larger work on the _Natural
History of Lichens_, but only two of the volumes reached publication.
In the first of these he devoted much attention to the “gonidia” or
“brood-cells” and established the distinction between the heteromerous
and homoiomerous distribution of green cells within the thallus; he also
describes with great detail the “morphosis” and “metamorphosis” of the
vegetative body. In the second volume he discusses their physiology—the
contents and products of the thallus, colouring, nutrition, season of
development, etc.—and finally the pathology of these organisms. He made
no great use of the compound microscope, and his studies were confined to
phenomena that could be observed with a single lens.
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