Thallus crustaceous squamulose.
Apothecia immersed in thalline warts 1. *Calothricopsis Wain.
Apothecia superficial, with thalline
margin 2. *Steinera A. Zahlbr.
Apothecia superficial, without a
thalline margin 3. Pterygium Nyl.
Thallus of small fruticose fronds.
Gonidia occupying the central strand 4. *Lichinodium Nyl.
Gonidia not in the centre.
Apothecia immersed 5. Lichina Ag.
Apothecia superficial.
Paraphyses present 6. *Lichinella Nyl.
Paraphyses absent 7. *Homopsella Nyl.
XXXIX. _COLLEMACEAE_
The most important family of the gelatinous lichens and the most
numerous. _Collema_ is historically interesting as having first suggested
the composite thallus. Algal cells, _Nostoc_, which retain the chain-like
form except in _Leprocollema_, a doubtful member of the family. The
thallus varies from indeterminate crusts to lobes of considerable size;
occasionally the lobes are narrow and erect, forming minute fruticose
structures. In the more primitive genera the thallus is non-corticate,
but in the more evolved, the apical cells of the hyphae coalesce to form
a continuous cellular cortex, one or more cells thick, well marked in
some species, in others rudimentary; the formation of plectenchyma also
occurs occasionally in the apothecial tissues of some non-corticate
species.
The apothecia are superficial except in _Pyrenocollema_, a monotypic
genus of unknown locality. They are generally lecanorine, with gonidia
entering into the formation of the apothecium: in some genera they are
lecideine or biatorine, being formed of hyphae alone. The spores are
colourless and vary in form, size and septation.
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