The difficulty of determining the primitive stations of cosmopolitan,
or of widely spread, species is much greater, but generally they also
may be referred to their area of greatest abundance. Thus a species
may occur frequently in one continent and but rarely in another, even
where the conditions of climate, etc., are largely comparable. It may
therefore be inferred that the plant has not yet reached the full extent
of possible distribution in the less frequented area. As examples of
this, Wainio cites, among other instances, _Cladonia papillaria_, which
has a very wide distribution in Europe, but, as yet, has been found only
in the eastern parts of North America; and _Cl. pycnoclada_, a plant
which braves the climate of Cape Horn and the Falkland Islands, but has
not travelled northward beyond temperate North America: the southern
origin of that species is thus plainly indicated. Wainio also finds that
evidence of the primitive locality of a very widely spread species may be
obtained by observing the locality of species derived from it, which are
as yet of limited distribution; presumably these arose in the ancestral
place of origin, though this indication is not always to be relied on.
If, however, the ancestral plant has given rise to several of these
rarer related species, those of them that are most closely allied to the
primitive plant would be found near to it in the original locality.
A detailed account of species distribution according to these indications
is given by Wainio and is full of interest. No such attempt has been
made to deal with any other group, and the distribution of genera and
species can only be suggested. An exhaustive comparison of the lichens
of different regions is beyond the purpose of our study and is indeed
impossible as, except in some limited areas, or for certain species,
the occurrence and distribution are not fully known. It is in any case
only tentatively that genera or species can be described as local or
rare, until diligent search has been made for them over a wider field.
The study of lichens from a floristic point of view lags behind that of
most other groups of plants. The larger lichen forms have received more
attention, as they are more evident and more easily collected; but the
more minute species are not easily detected, and, as they are largely
inseparable from their substratum of rocks, or trees, etc., on which
they grow, they are often difficult to collect. They are also in many
instances so indefinite, or so alike in outward form, that they are
liable to be overlooked, only a microscopic examination revealing the
differences in fruit and vegetative structure.
Though much remains to be done, still enough is known to make the
geographical distribution of lichens a subject of extreme interest. It
will be found most instructive to follow the usual lines of treatment,
which give the three great divisions: the Polar, the Temperate and the
Tropical regions of the globe.
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