B. LICHENS OF POLAR REGIONS
Strictly speaking, this section should include only lichens growing
within the Polar Circles; but in practice the lichens of the whole of
Greenland and those of Iceland are included in the Arctic series, as
are those of Alaska: the latitudinal line of demarcation is not closely
adhered to. With the northern lichens may also be considered those of the
Antarctic continent, as well as those of the islands just outside the
Antarctic Circle, the South Shetlands, South Orkneys, Tierra del Fuego,
South Georgia and the Falkland Islands. During the Glacial period, the
polar forms must have spread with the advancing cold; as the snow and ice
retreated, these forms have been left, as already stated, on the higher
colder grounds, and representatives of polar species are thus to be found
very far from their original haunts. There are few exclusively boreal
genera: the same types occur at the Poles as in the higher subtemperate
zones. One of the most definitely polar species, for instance, _Usnea_
(_Neuropogon_) _melaxantha_ grows in the whole Arctic zone, and, in the
Antarctic, is more luxuriant than any other lichen, but it has also been
recorded from the Andes in Chili, Bolivia and Peru, and from New Zealand
(South Island).
Cold winds are a great feature of both poles, and the lichens that by
structure or habit can withstand these are the most numerous; those that
have a stout cortical layer are able to resist the low temperatures, or
those that grow in tufts and thus secure mutual protection. In Arctic
and Subarctic regions, 495 lichens have been recorded, most of them
crustaceous. Among the larger forms the most frequently met are certain
species of _Peltigera_, _Parmelia_, _Gyrophora_, _Cetraria_, _Cladonia_,
_Stereocaulon_ and _Alectoria_. Among smaller species _Lecanora tartarea_
spreads everywhere, especially over other vegetation, _Lecanora varia_
reaches the farthest limits to which wood, on which it grows, has
drifted, and several species of _Placodium_ occur constantly, though not
in such great abundance. Over the rocks spread also many crustaceous
Lecideaceae too numerous to mention, one of the most striking being the
cosmopolitan _Rhizocarpon geographicum_.
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