Life and Letters of Charles Darwin — Volume 2Darwin, Charles
Science
Life and Letters of Charles Darwin — Volume 2
Darwin, Charles
Darwin, Charles, 1809-1882; Naturalists -- Great Britain -- Biography; Naturalists -- Great Britain -- Correspondence
On the theory of Natural Selection there is a wide distinction between
Rudimentary Organs and what you call germs of organs, and what I call
in my bigger book "nascent" organs. An organ should not be called
rudimentary unless it be useless--as teeth which never cut through the
gums--the papillae, representing the pistil in male flowers, wing of
Apteryx, or better, the little wings under soldered elytra. These organs
are now plainly useless, and a fortiori, they would be useless in a
less developed state. Natural Selection acts exclusively by preserving
successive slight, USEFUL modifications. Hence Natural Selection cannot
possibly make a useless or rudimentary organ. Such organs are solely due
to inheritance (as explained in my discussion), and plainly bespeak an
ancestor having the organ in a useful condition. They may be, and
often have been, worked in for other purposes, and then they are only
rudimentary for the original function, which is sometimes plainly
apparent. A nascent organ, though little developed, as it has to be
developed must be useful in every stage of development. As we cannot
prophesy, we cannot tell what organs are now nascent; and nascent organs
will rarely have been handed down by certain members of a class from a
remote period to the present day, for beings with any important organ
but little developed, will generally have been supplanted by their
descendants with the organ well developed. The mammary glands in
Ornithorhynchus may, perhaps, be considered as nascent compared with
the udders of a cow--Ovigerous frena, in certain cirripedes, are nascent
branchiae--in [illegible] the swim bladder is almost rudimentary for
this purpose, and is nascent as a lung. The small wing of penguin, used
only as a fin, might be nascent as a wing; not that I think so; for
the whole structure of the bird is adapted for flight, and a penguin
so closely resembles other birds, that we may infer that its wings have
probably been modified, and reduced by natural selection, in accordance
with its sub-aquatic habits. Analogy thus often serves as a guide in
distinguishing whether an organ is rudimentary or nascent. I believe the
Os coccyx gives attachment to certain muscles, but I can not doubt that
it is a rudimentary tail. The bastard wing of birds is a rudimentary
digit; and I believe that if fossil birds are found very low down in the
series, they will be seen to have a double or bifurcated wing. Here is a
bold prophecy!
To admit prophetic germs, is tantamount to rejecting the theory of
Natural Selection.
Public-domain text, read in full here on John Shaqi.
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