Life Movements in Plants, Volume IBose, Jagadis Chandra
Science
Life Movements in Plants, Volume I
Bose, Jagadis Chandra
Growth (Plants); Plants -- Irritability and movements
I will first give a diagrammatic view of the different parts of the
apparatus which I devised for this investigation.[G] The leaf of
_Mimosa_ is attached to one arm of a light aluminium lever, L, by means
of thread. At right angles to the lever is the writing index W, which
traces on a smoked glass plate allowed to fall at a definite rate
by clockwork the responsive movement of the leaf. Under a definite
stimulus of electric shock the leaf falls down, pulling the lever L,
and moving the writer towards the left. (Fig. 14.) The amplitude of
the response-curve measures the intensity of excitation. The leaf
re-erects itself after a time, the corresponding record exhibiting
recovery. A second stimulus is applied after a definite interval, say
an hour, and the corresponding response shows whether the excitability
of the plant has remained constant or undergone any variation.
[G] _See_ also Bose.--The Diurnal Variation of Moto-excitability in
_Mimosa_--Annals of Botany, Oct. 1913.
[Illustration: FIG. 14. Diagrammatic representation of the complete
apparatus for determination of diurnal variation of excitability.
Petiole of _Mimosa_, attached by thread to one arm of lever L;
writing index W traces on smoked glass plate G, the responsive fall
and recovery of leaf. A, primary, and S, secondary, of induction
coil. Exciting shock passes through the plant by electrodes E, E′. A,
accumulator. C, clockwork for regulating duration of tetanizing shock.
Primary circuit of coil completed by plunging rod, V, dipping into cup
of mercury M.]
UNIFORM PERIODIC STIMULATION.
_Electric mode of excitation._--I find that one of the best methods of
stimulating the plant is by means of tetanizing induction shock. The
sensitiveness of _Mimosa_ to electric stimulation is very great; the
plant often responds to a shock which is quite imperceptible to a human
subject. By the employment of a sliding induction coil, the intensity
of the shock can be regulated with great accuracy; the secondary if
gradually brought nearer the primary till a stimulus is found which is
minimally effective. The intensity of stimulus actually employed is
slightly higher than this, but within the sub-maximal range. When the
testing stimulus is maintained constant and of sub-maximal intensity,
then any variation of excitability is attended by a corresponding
variation in the amplitude of response.
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