Life Movements in Plants, Volume IBose, Jagadis Chandra
Science
Life Movements in Plants, Volume I
Bose, Jagadis Chandra
Growth (Plants); Plants -- Irritability and movements
The Oscillator is diagrammatically shown in Fig. 15. M, M′ are the two
electromagnetic coils, the free ends of the horseshoe being pointed.
Facing them are the conical holes of the soft iron armature A. This
armature carries two rods which slide through hollow tubes. The distal
ends of the rods support the holder H, carrying the smoked glass plate.
Under normal conditions, the plate-holder is held by suitable springs,
somewhat to the right of, and free from contact with, the writer. A
clockwork C carries a rotating arm, which makes periodic contact with
a pool of mercury contained in the vessel V, once in a minute. On the
completion of the electromagnetic circuit, the armature A is attracted,
the recording glass plate being thereby moved to the left making
contact with the writer. The successive dots in the record thus take
place at intervals of a minute. Only a moderate amount of electric
current is thus consumed in maintaining the oscillation of the plate. A
4-volt storage cell of 20 amperes capacity is quite sufficient to work
the apparatus for several days.
The responsive fall of the leaf of _Mimosa_ is completed in the course
of about two seconds. The leaf remains in the fallen or ‘contracted’
position for nearly fifteen seconds; it then begins to recover slowly.
As the successive dots of the Oscillating Recorder are at intervals
of a minute, the maximum fall of leaf is accomplished between two
successive dots. The dotted response record here obtained exhibits
the recovery from maximum fall under stimulation (_cf._ Fig. 23). The
recovery of the leaf in one minute is less than one-tenth the total
amplitude of the fall, and is proportionately the same in all the
response records. Hence the successive amplitudes of response curves
that are recorded at different hours of the day afford us measures
of the relative variations of excitability of the plant at different
times. This enables us to demonstrate the reality of diurnal variation
of excitability. In my experimental investigations on the subject I
have not been content to take my data from any particular method of
obtaining response, but have employed both types of recorders, the
Resonant and Oscillating. It will be shown that the results given by
the different instruments are in complete agreement with each other.
EFFECTS OF EXTERNAL CONDITIONS ON EXCITABILITY.
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