Life Movements in Plants, Volume IIBose, Jagadis Chandra
Science
Life Movements in Plants, Volume II
Bose, Jagadis Chandra
Growth (Plants); Plants -- Irritability and movements
In the former case the upper half of the pulvinus occupied the
up-position; in the second case the up-position was occupied by the
lower half of the pulvinus. In both cases strong electric responses were
obtained, the upper point of contact being always galvanometrically
negative. There was, however, a difference between the two responses,
the excitatory electro-motive variation was invariably greater when the
lower half of the organ occupied the favourable up-position. This will
be seen from the results of two typical experiments in table given
above.
The electrical mode of investigation thus leads to confirm the result
obtained with mechanical method that the lower half of the pulvinus of
_Mimosa_ is geotropically more excitable than the upper half.
RELATION BETWEEN ANGLE OF INCLINATION AND GEOTROPIC EFFECT.
In the Method of Axial Rotation, the condition of the experiment is
ideally perfect; in the neutral position the sides A and B are both
parallel to the vertical lines of gravity, and are little affected by
geotropic reaction. As the specimen is rotated on its long axis the
vertical component of the force of gravity increases with the angle of
inclination. The hypothetical statolithic particles will become
displaced all along the cell, and the vertical pressure exerted by them
will also increase with the angle.
The geo-electric response will then afford us a measure of the intensity
of excitation induced at various angles of inclination. The mechanical
response on account of its inherent defects does not afford us the true
relation between the angle of inclination and intensity of geotropic
reaction. But the electric method of inquiry is free from the defects of
the mechanical method.
_Experiment 176._--The specimen was rotated so that the angle of
rotation was 45°, and the maximum electric response observed. The angle
was next increased to 90° and the reading for the enhanced response
taken. The ratio of the geo-electric response at 90° and 45°, thus
affords us a measure of the effective stimulations at the two angles.
I give below a table which gives results obtained with 24 different
specimens.
TABLE XXXV.--RELATION BETWEEN ANGLE OF INCLINATION AND GEOTROPIC EFFECT.
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