Life Movements in Plants, Volume IIBose, Jagadis Chandra
Science
Life Movements in Plants, Volume II
Bose, Jagadis Chandra
Growth (Plants); Plants -- Irritability and movements
For purpose of simplicity of explanation, I assumed the growth rate to
have a definite value of 1/50,000 inch per second. But the rate varies
widely in different plants and even in the same plant at different days
and seasons. In practice the rate of growth for which compensation has
to be made varies from 1/150,000 to 1/25,000 inch, or from 0·17 µ to
1·0 µ per second. We have thus to secure some means of _continuous_
adjustment for growth, the rate of which could be continuously varied
from one to six times. This range of adjustment I have been able to
secure by the compound method of frictional resistance and of
centrifugal governor. As regards frictional resistance the two pointed
ends of a hinged fork rub against a horizontal circular plate not shown
in the figure. By means of the screw head S, the free ends of the fork
spread out and the circumference of the frictional circle continuously
increased. The centrifugal governor is also spread out by the action of
the adjusting screw. By the joint actions of the frictional control and
the centrifugal governor, the speed of rotation can be continuously
adjusted from 1 to 6 times. When the adjusting screw is set in a
particular position, the speed of rotation, and therefore the rate of
subsidence of plant-holder, remains absolutely constant for several
hours. The attainment of this constancy is a matter of fundamental
importance, and it was only by the employment or the compound system of
regulation that I was able to secure it.
The method of obtaining balance now becomes extremely simple. Before
starting the balancing movement by clock regulation, the plant is made
to record its magnified growth by the Crescograph. The compensation is
effected as follows: the speed of the clockwork is at the beginning
adjusted at its lowest value, and the pressure of a button starts the
balancing movement of the plant downwards. On account of partial balance
the record will be found to be less steep than before; the speed of the
clock is gradually increased till the record becomes perfectly
horizontal under exact balance. Overbalance makes the record slope
downwards. In figure 95 is seen records of underbalance (_a_) and
overbalance (_b_), to the extent of about 3 per cent.
[Illustration: FIG. 95.--Balanced Crescographic record: (_a_) showing
effect of underbalance and (_b_) overbalance of about 3 per cent.
(Magnification 2,000 times.)]
It will thus be seen that the effect of an external agent may be
detected by the upsetting of the balance; an up-movement indicates
(unless stated to the contrary) an enhancement of the rate of growth
above the normal; and a down-movement, on the other hand, a depression
of the normal rate.
Public-domain text, read in full here on John Shaqi.
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