The experiments that have just been described are of much significance. An organ like the stem of _Mimosa_, since it exhibits no contraction, may appear insensitive to stimulation; but its perception of stimulus is shown by its power of transmitting two characteristic impulses, one of which is the positive, giving rise to an enhancement of turgor, and the other, the true excitatory negative, inducing the opposite reaction or diminution of turgor. Unilateral stimulation gives rise to both these effects in all organs: pulvinated, growing, and non-growing. It was the fortunate circumstance of the insertion of the motile leaf on one side of the _Mimosa_ stem that enabled us to demonstrate the important facts given above. The underlying reactions, which give rise to tropic curvature, could have been foretold from the Laws of effects of Direct and Indirect stimulation, established in previous chapters (pp. 136, 216). The resulting curvature is thus brought about by the joint effects of direct stimulation of the proximal, and indirect stimulation of the distal side. We may now recapitulate some of the important facts relating to tropic curvatures: Indirect stimulation gives rise to dual impulses, positive and negative; of these the positive impulse is practically independent of the conducting power of the tissue; but the transmission of the excitatory negative impulse is dependent on the conducting power. No tissue is a perfect conductor, nor is any a perfect non-conductor of excitation, the difference is a question of degree. In a petiole or a stem the conducting power along the direction of length is considerable, but very feeble in a transverse direction. In a semi-conducting tissue, a feeble stimulus will transmit only the positive impulse; strong or long continued stimulation will transmit both positive and negative impulses, the positive preceding the negative. The transmitted positive gives rise to increase of turgor, expansion, and acceleration of rate of growth; the negative induces the opposite reaction of diminution of turgor, of contraction, and of retardation of rate of growth. Transverse transmission is only a particular instance of transmission in general; the only difference is that the conducting power for _excitation_ is very much less in the transverse than in the longitudinal direction. Owing to feeble transverse conductivity, the transmitted impulse to the distal side often remains positive; it is only under strong or continued stimulation that the excitatory negative reaches the distal side and neutralises or reverses the previous positive reaction. If the distal is the more excitable side, the reversed response will appear as pronounced negative. I give a table which will clearly exhibit the effects of stimulus on the proximal and distal sides of the responding organ. TABLE XXIV.--SHOWING RESPONSIVE EFFECTS COMMON TO PULVINI AND GROWING ORGANS UNDER UNILATERAL STIMULATION.