In the Iberian peninsula the tundra fauna did not penetrate as far south as Portugal, although the Norwegian lemming (_Myodes lemmus_) reached the vicinity of Lisbon. The woolly mammoth, accompanied by the woolly rhinoceros, has been discovered in two localities on the extreme northern coast of Spain, in the province of Santander, bordering the Bay of Biscay. The reindeer (_Rangifer tarandus_) is found in the cavern of Seriña, south of the Pyrenees; as early as Acheulean times it reached the region of Altamira, near Santander. Thus Harlé(24) concludes it is certain that the tundra fauna spread from France westward into Catalonia, along the northern coast of Spain, flanking the Pyrenees. It is generally believed that the cave-bear (_Ursus spelæus_) occupied many of the caverns before their possession by man, and developed certain peculiarities of structure in these haunts. Thus the phalanges bearing the claws are feebly developed, indicating that the claws had partly lost their prehensile function; the anterior grinding-teeth are very much reduced, and the cusps of the posterior grinders are blunted in a way which is indicative of an omnivorous diet; yet the front paws were of tremendous size, the body was thick-set and of heavier proportions than that of the larger recent bears (_Ursus arctos_) of Europe. Hence, it would appear that the Neanderthals drove out from the caves a type of bear less formidable than the existing species but nevertheless a serious opponent to men armed with the small weapons of the Mousterian period. CUSTOMS OF THE CHASE AND OF CAVE LIFE We have only indirect means of knowing the courage and activity of the Neanderthals in the chase, through the bones of animals hunted for food which are found intermingled with the flints around their ancient hearths. These include in the early Mousterian hearths, as we have seen, bones of the bison, the wild cattle, and the horse, which are followed at Combe-Capelle by the first appearance of the bones of the reindeer. The bones of the bison and of the wild horse are both utilized in the bone anvils of the closing Mousterian culture at La Quina. What we believe to be the period of the great mammalian life of the region of the upper Danube is found in the Mousterian levels of the grotto of Sirgenstein, from which it would appear that the Neanderthals hunted the mammoth, the rhinoceros, the wild horse, bison, and cattle, and the giant deer as well as the reindeer. We should keep in mind, however, that when these caves were for a time deserted, the beasts of prey returned, and so it often happens that the succeeding layers afford proofs of alternate occupation by man and by beasts of prey of sufficient size to bring in the larger kinds of game, while owls may be responsible for the deposits of the lemming, as in the 'lower rodent layer.'