Mendel's principles of heredity: A defenceMendel, Gregor
Science
Mendel's principles of heredity: A defence
Mendel, Gregor
Heredity; Hybridization; Mendel's law; Weldon, Walter Frank Raphael. Mendel's laws of alternative inheritance in peas
To go into the whole question in detail would require considerable
space; but indeed it is unnecessary to labour the point. The reader who
examines Gärtner’s account with care, especially the peculiar phenomena
obtained in the case of the “grey” pea (_macrospermum_), with specimens
before him, will have no difficulty in recognizing that Gärtner is
simply describing the seeds _as they looked in their coats_, and is not
attempting to distinguish cotyledon-characters and coat-characters.
If he had peeled them, which in the case of “grey” peas would be
_absolutely necessary_ to see cotyledon-colour, he must surely have
said so.
Had he done so, he would have found the cotyledons full yellow in every
ripe seed; for I venture to assert that anyone who tries, as we have,
crosses between a yellow-cotyledoned “grey” pea, such as Gärtner’s was,
with any pure green variety will see that there is no question whatever
as to absolute dominance of the yellow cotyledon-character here, more
striking than in any other case. If exceptions are to be looked for,
they will not be found _there_; and, except in so far as they show
simple dominance of yellow, Gärtner’s observations cannot be cited in
this connection at all.
(2) _Seton’s case._ Another exception given by Professor Weldon is much
more interesting and instructive. It is the curious case of Seton[86].
Told in the words of the critic it is as follows:--
“Mr Alexander Seton crossed the flowers of _Dwarf Imperial_, ‘a
well-known green variety of the Pea,’ with the pollen of ‘a white
free-growing variety.’ Four hybrid seeds were obtained, ‘which did
not differ in appearance from the others of the female parent.’
These seeds therefore did _not_ obey the law of dominance, or if the
statement be preferred, greenness became dominant in this case. The
seeds were sown, and produced plants bearing ‘green’ and ‘white’
seeds side by side in the same pod. An excellent coloured figure of
one of these pods is given (_loc. cit._ Plate 9, Fig. 1), and is the
only figure I have found which illustrates segregation of colours in
hybrid Peas of the second generation.”
[86] Appendix to paper of Goss, _Trans. Hort. Soc._ v. 1822, pub.
1824 (_not_ 1848, as given by Professor Weldon), p. 236.
Now if Professor Weldon had applied to this case the same independence
of judgment he evinced in dismissing Darwin’s interpretation of
Gärtner’s observations, he might have reached a valuable result.
Knowing how difficult it is to give all the points in a brief citation,
I turned up the original passage, where I find it stated that the mixed
seeds of the second generation “were all completely either of one
colour or the other, none of them having an intermediate tint, as Mr
Seton had expected.” The utility of this observation of the absence of
intermediates, is that it goes some way to dispose of the suggestion of
xenia as a cause contributing to the result.
Public-domain text, read in full here on John Shaqi.
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