Mendel's principles of heredity: A defenceMendel, Gregor
Science
Mendel's principles of heredity: A defence
Mendel, Gregor
Heredity; Hybridization; Mendel's law; Weldon, Walter Frank Raphael. Mendel's laws of alternative inheritance in peas
Moreover, feeling perfectly clear, from the fact of the absence of
intermediates, that the case must be one of simple dominance in
spite of first appearances, I suggest the following account with
every confidence that it is the true one. There have been several
“_Imperials_,” though _Dwarf Imperial_, in a form which I can feel
sure is Seton’s form, I have not succeeded in seeing; but from
Vilmorin’s description that the peas when ripe are “_franchement
verts_” I feel no doubt it was a green pea _with a green skin_. If it
had had a transparent skin this description would be inapplicable.
Having then a green skin, which may be assumed with every probability
of truth, the seeds, even though the cotyledons were yellow, might,
especially if examined fresh, be indistinguishable from those of
the maternal type. Next from the fact of the mixture in the second
generation we learn that the _semi-transparent seed-coat of the
paternal form was dominant_ as a plant-character, and indeed the
coloured plate makes this fairly evident. It will be understood that
this explanation is as yet suggestive, but from the facts of the
second generation, any supposition that there was real irregularity in
dominance in this case is out of the question[87].
[87] Since the above passage was written I find the “_Imperials_”
described in “Report of Chiswick Trials,” _Proc. R. Hort. Soc._ 1860,
I. p. 340, as “skin thick”; and on p. 360 “skin thick, blue”; which
finally disposes of this “exception.”
(3) _Tschermak’s exceptions._ These are a much more acceptable lot
than those we have been considering. Tschermak was thoroughly alive
to the seed-coat question and consequently any exception stated as an
unqualified fact on his authority must be accepted. The nature of these
cases we shall see. Among the many varieties he used, some being _not_
monomorphic, it would have been surprising if he had not found true
irregularities in dominance.
(3 _a_) _Buchsbaum case._ This variety, growing in the open, gave
once a pod in which _every seed but one was green_. In stating this
case Professor Weldon refers to _Buchsbaum_ as “a yellow-seeded
variety.” Tschermak[88], however, describes it as having “_gelbes,
öfters gelblich-grünes Speichergewebe_” (cotyledons); and again
says the cotyledon-colour is “_allerdings gerade bei Buchsbaum zur
Spontanvariation nach gelb-grün neigend!_” The (!) is Tschermak’s.
Therefore Professor Weldon can hardly claim _Buchsbaum_ as
“yellow-seeded” without qualification.
[88] (36), p. 502 and (37), p. 663.
_Buchsbaum_ in fact is in all probability a blend-form and certainly
not a true, stable yellow. One of the green seeds mentioned above grew
and gave 15 _yellows_ and three _greens_, and the result showed pretty
clearly, as Tschermak says, that there had been an accidental cross
with a tall green.
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