Mendel's principles of heredity: A defenceMendel, Gregor
Science
Mendel's principles of heredity: A defence
Mendel, Gregor
Heredity; Hybridization; Mendel's law; Weldon, Walter Frank Raphael. Mendel's laws of alternative inheritance in peas
“So the cross _Couturier_ (orange-yellow) ♀ × the green-seeded
_Express_ ♂ gave a number of seeds intermediate in colour. (It is not
clear from Tschermak’s paper whether _all_ the seeds were of this
colour, but certainly some of them were.) The green _Plein le Panier_
[_Fillbasket_] ♀ × _Couturier_ ♂ in three crosses always gave either
seeds of colour intermediate between green and yellow, or some yellow
and some green seeds in the same pod. The cross reciprocal to this was
not made; but _Express_ ♀ × _Couturier_ ♂ gave 22 seeds of which four
were yellowish green[149].
[149] See p. 148.
“These facts show _first_ that Mendel’s law of dominance conspicuously
fails for crosses between certain races, while it appears to hold
for others; and _secondly_ that the intensity of a character in one
generation of a race is no trustworthy measure of its dominance in
hybrids. The obvious suggestion is that the behaviour of an individual
when crossed depends largely upon the characters of its ancestors[150].
When it is remembered that peas are normally self-fertilised, and that
more than one named variety may be selected out of the seeds of a
single hybrid pod, it is seen to be probable that Mendel worked with a
very definite combination of ancestral characters, and had no proper
basis for generalisation about yellow and green peas of any ancestry”
[which he never made].
[150] Where was that “logician,” the “consulting-partner,” when this
piece of reasoning passed the firm?
Let us pause a moment before proceeding to the climax. Let the reader
note we have been told of _two_ groups of cases in which dominance of
yellow failed or was irregular. (Why are not Gärtner’s and Seton’s
“exceptions” referred to here?) In one of these groups _Couturier_
was always one parent, either father or mother, and were it not for
Tschermak’s own obvious hesitation in regard to his own exceptions
(see p. 148), I would gladly believe that _Couturier_--a form I do not
know--may be an exceptional variety. _How_ Professor Weldon proposes
to explain its peculiarities by reference to ancestry he omits to tell
us. The _Buchsbaum_ case is already disposed of, for on Tschermak’s
showing, it is an unstable form.
Public-domain text, read in full here on John Shaqi.
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