Mendel's principles of heredity: A defenceMendel, Gregor
Science
Mendel's principles of heredity: A defence
Mendel, Gregor
Heredity; Hybridization; Mendel's law; Weldon, Walter Frank Raphael. Mendel's laws of alternative inheritance in peas
In view of the evidence, the conclusion is forced upon me that the
suggestion that “ancestry” may explain the facts regarding _Telephone_
has no meaning behind it, but is merely a verbal obstacle. Two words
more on _Telephone_. On p. 147 I ventured to hint that if we try to
understand the nature of the appearance of green in the offspring
of _Telephone_ bred with yellow varieties, we are more likely to do
so by comparing the facts with those of false hybridisation than
with fluctuations in dominance. In this connection I would call the
reader’s attention to a point Professor Weldon misses, that Tschermak
_also got yellowish-green seeds from Fillbasket (green) crossed with
Telephone_. I suggest therefore that _Telephone’s_ allelomorphs may be
in part transmitted to its offspring in a state which needs no union
with any corresponding allelomorph of the other gamete, just as may
the allelomorphs of “false hybrids.” It would be quite out of place
here to pursue this reasoning, but the reader acquainted with special
phenomena of heredity will probably be able fruitfully to extend it.
It will be remembered that we have already seen the further fact that
the behaviour of _Telephone_ in respect to seed-shape was also peculiar
(see p. 152).
Whatever the future may decide on this interesting question it is
evident that with _Telephone_ (and possibly _Buchsbaum_) we are
encountering a _specific_ phenomenon, which calls for specific
elucidation and not a case simply comparable with or contradicting the
evidence of dominance in general.
In this excursion we have seen something more of the “exceptions.”
Many have fallen, but some still stand, though even as to part of
the remainder Tschermak entertains some doubts, and, it will be
remembered, cautions his reader that of his exceptions some may be
self-fertilisations, and some did not germinate[152]. Truly a slender
basis to carry the coming structure!
[152] In his latest publication on this subject, the notes to the
edition of Mendel in Ostwald’s _Klassiker_ (pp. 60–61), Tschermak,
who has seen more true exceptions than any other observer, thus
refers to them. As to dominance:--“_Immerhin kommen vereinzelt
auch zweifellose Fälle von Merkmalmischung, d. h. Uebergangsformen
zwischen gelber und grüner Farbe, runder und runzeliger Form vor,
die sich in weiteren Generationen wie dominantmerkmalige Mischlinge
verhalten._” As to purity of the extracted recessives:--_Ganz
vereinzelt scheinen Ausnahmsfälle vorzukommen._"
Küster (22) also in a recent note on Mendelism points out, with
reason, that the number of “exceptions” to dominance that we shall
find, depends simply on the stringency with which the supposed “law”
is drawn. The same writer remarks further that Mendel makes no such
rigid definition of dominance as his followers have done.
Public-domain text, read in full here on John Shaqi.
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