Mendel's principles of heredity: A defenceMendel, Gregor
Science
Mendel's principles of heredity: A defence
Mendel, Gregor
Heredity; Hybridization; Mendel's law; Weldon, Walter Frank Raphael. Mendel's laws of alternative inheritance in peas
Generally speaking peas which when seen in bulk are monomorphic in
colour and shape, will give fairly true and uniform offspring (but
such strict monomorphism is rather exceptional). Instances to the
contrary occur, and in my own brief experience I have seen some.
In a row of _Fill-basket_ grown from selected seed there were two
plants of different habit, seed-shape, etc. Each bore pods with seeds
few though large and round. Again _Blue Peter_ (blue and round) and
_Laxton’s Alpha_ (blue and wrinkled), grown in my garden and left to
nature uncovered, have each given a considerable proportion of seeds
with _yellow_ cotyledons, about 20% in the case of _Laxton’s Alpha_.
The distribution of these on the plants I cannot state. The plants
bearing them in each case sprang from green-cotyledoned seeds taken
from samples containing presumably unselected green seeds only. A part
of this exceptional result may be due to crossing, but heterogeneity of
conditions[68] especially in or after ripening is a more likely cause,
hypotheses I hope to investigate next season. Hitherto I had supposed
the crossing, if any, to be done by _Bruchus_ or Thrips, but Tschermak
also suspects _Megachile_, the leaf-cutter bee, which abounds in my
garden.
[68] Cotyledon-colour is not nearly so sensitive to ordinary changes
in conditions as coat-colour, provided the coat be uninjured. But
even in monomorphic _green_ varieties, a seed which for any cause has
burst on ripening, has the exposed parts of its cotyledons _yellow_.
The same may be the case in seeds of green varieties injured by
_Bruchus_ or birds. These facts make one hesitate before denying
the effects of conditions on the cotyledon-colour even of uninjured
seeds, and the variation described above may have been simply
weathering. The seeds were gathered very late and many were burst in
_Laxton’s Alpha_. I do not yet know they are alive.
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