Mendel's principles of heredity: A defenceMendel, Gregor
Science
Mendel's principles of heredity: A defence
Mendel, Gregor
Heredity; Hybridization; Mendel's law; Weldon, Walter Frank Raphael. Mendel's laws of alternative inheritance in peas
Whatever the cause, these irregularities may undoubtedly occur; and if
they be proved to be largely independent of crossing and conditions,
this will in nowise vitiate the truth of the Mendelian principle.
For in that case it may simply be variability. Such true variation,
or sporting, in the pea is referred to by many observers. Upon this
subject I have received most valuable facts from Mr Arthur Sutton,
who has very kindly interested himself in these inquiries. He tells
me that several highly bred varieties, selected with every possible
care, commonly throw a small but constant proportion of poor and almost
vetch-like plants, with short pods and small round seeds, which are
hoed out by experienced men each year before ripening. Other high-class
varieties always, wherever grown, and when far from other sorts,
produce a small percentage of some one or more definite “sports.” Of
these peculiar sports he has sent me a collection of twelve, taken from
as many standard varieties, each “sport” being represented by eight
seeds, which though quite distinct from the type agree with each other
in almost all cases.
In two cases, he tells me, these seed-sports sown separately have
been found to give plants identical with the standard type and must
therefore be regarded as sports in _seed characters_ only; in other
cases change of plant-type is associated with the change of seed-type.
In most standard varieties these definite sports are not very common,
but in a few they are common enough to require continual removal by
selection[69].
[69] It is interesting to see that in at least one case the same--or
practically the same--variety has been independently produced by
different raisers, as we now perceive, by the fortuitous combination
of similar allelomorphs. _Sutton’s Ringleader_ and _Carter’s First
Crop_ (and two others) are cases in point, and it is peculiarly
instructive to see that in the discussion of these varieties when
they were new, one of the points indicating their identity was taken
to be the fact that they produced _the same “rogues.”_ See _Gard.
Chron._ 1865, pp. 482 and 603; 1866, p. 221; 1867, pp. 546 and 712.
Rimpau quotes Blomeyer (_Kultur der Landw. Nutzpflanzen_, Leipzig,
1889, pp. 357 and 380) to the effect that _purple_-flowered plants
with _wrinkled_ seeds may spring as direct sports from peas with
_white_ flowers and _round_ seeds. I have not seen a copy of
Blomeyer’s work. Probably this “wrinkling” was “indentation.”
I hope before long to be able to give statistical details and
experiments relating to this extraordinarily interesting subject. As
de Vries writes in his fine work _Die Mutationstheorie_ (I. p. 580),
“a study of the seed-differences of inconstant, or as they are
called, ‘still’ unfixed varieties, is a perfect treasure-house of new
discoveries.”
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