Mendelism: Third EditionPunnett, Reginald Crundall
Science
Mendelism: Third Edition
Punnett, Reginald Crundall
Mendel's law
The task that Mendel set before himself was to gain some clear conception
of the manner in which the definite and fixed varieties found within a
species are related to one another, and he realised at the outset that the
best chance of success lay in working with material of such a nature as to
reduce the problem to its simplest terms. He decided that the plant with
which he was to work must be normally self-fertilising and unlikely to be
crossed through the interference of insects, while at the same time it must
possess definite fixed varieties which bred true to type. In the common pea
(_Pisum sativum_) he found the plant he sought. A hardy annual, prolific,
easily worked, _Pisum_ has a further advantage in that the insects which
normally visit flowers are unable to gather pollen from it and so to bring
about cross fertilisation. At the same time it exists in a number of
strains presenting well-marked and fixed differences. The flowers may be
purple, or red, or white; the plants may be tall or dwarf; the ripe seeds
may be yellow or green, round or wrinkled--such are a few of the characters
in which the various races of peas differ from one another. {18}
In planning his crossing experiments Mendel adopted an attitude which
marked him off sharply from the earlier hybridisers. He realised that their
failure to elucidate any general principle of heredity from the results of
cross fertilisation was due to their not having concentrated upon
particular characters or traced them carefully through a sequence of
generations. That source of failure he was careful to avoid, and throughout
his experiments he crossed plants presenting sharply contrasted characters,
and devoted his efforts to observing the behaviour of these characters in
successive generations. Thus in one series of experiments he concentrated
his attention on the transmission of the characters tallness and dwarfness,
neglecting in so far as these experiments were concerned any other
characters in which the parent plants might differ from one another. For
this purpose he chose two strains of peas, one of about 6 feet in height,
and another of about 1-1/2 feet. Previous testing had shown that each
strain bred true to its peculiar height. These two strains were
artificially crossed[1] with one another, and it was found to make no
difference which was used as the pollen parent and which was used as the
ovule parent. In either case the result was the same. The result of
crossing tall with dwarf was in every case nothing but talls, as tall or
even a little taller than the tall parent. For this reason Mendel termed
tallness the DOMINANT and {19} dwarfness the RECESSIVE character. The next
stage was to collect and sow the seeds of these tall hybrids. Such seeds in
the following year gave rise to a mixed generation consisting of talls and
dwarfs _but no intermediates_. By raising a considerable number of such
plants Mendel was able to establish the fact that the number of talls which
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