Mendelism: Third EditionPunnett, Reginald Crundall
Science
Mendelism: Third Edition
Punnett, Reginald Crundall
Mendel's law
{81} Again, the proportion in which the three classes of purples appeared
was 9 bicolors, 3 deep purples, 4 picotees. We are, therefore, concerned
here with the operation of two factors: (1) a light wing factor, which
renders the bicolor dominant to the dark winged form; and (2) a factor for
intense colour, which occurs in the bicolor and in the deep purple, but is
lacking in the dilute picotee. And here it should be mentioned that these
conclusions rest upon an exhaustive set of experiments involving the
breeding of many thousands of plants. In this cross, therefore, we are
concerned with the presence or absence of five factors, which we may denote
as follows:--
A colour base, R.
A colour developer, C.
A purple factor, B.
A light wing factor, L.
A factor for intense colour, I.
On this notation our six coloured forms are:--
(1) Purple bicolor CRBLI.[6]
(2) Deep purple CRBlI.
(3) Picotee CRBLi or CRBli.
(4) Red bicolor ( = Painted Lady) CRbLI.
(5) Deep red ( = Miss Hunt) CRblI.
(6) Tinged white CRbLi or CRbli.
It will be noticed in this series that the various coloured {82} forms can
be expressed by the omission of one or more factors from the purple bicolor
of the wild type. With the complete omission of each factor a new colour
type results, and it is difficult to resist the inference that the various
cultivated forms of the sweet pea have arisen from the wild by some process
of this kind. Such a view tallies with what we know of the behaviour of the
wild form when crossed by any of the garden varieties. Wherever such
crossing has been made the form of the hybrid has been that of the wild,
thus supporting the view that the wild contains a complete set of all the
differentiating factors which are to be found in the sweet pea.
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