Mendelism: Third EditionPunnett, Reginald Crundall
Science
Mendelism: Third Edition
Punnett, Reginald Crundall
Mendel's law
Here, again, the calculated results accord
with those of experiment. Lastly, we may examine what should happen when
the F_1 female is crossed with the _lacticolor_ {104} male. The F_1 female,
owing to the repulsion between F and G, produces only the two kinds of ova
Fg and fG, and produces them in equal numbers. Since the _lacticolor_ male
can contain neither F nor G, all of its spermatozoa must be fg. The results
of such a cross, therefore, should be to produce equal numbers of the two
kinds of zygote Ffgg and ffGg, _i.e._ of _lacticolor_ females and of
_grossulariata_ males. And this, as we have already seen, is the actual
result of such a cross.
Before leaving the currant moth we may allude to an interesting discovery
which arose out of these experiments. The _lacticolor_ variety in Great
Britain is a southern form and is not known to occur in Scotland. Matings
were made between wild Scotch females and _lacticolor_ males. The families
resulting from such matings were precisely the same as those from
_lacticolor_ males and F_1 females, viz. _grossulariata_ males and
_lacticolor_ females only. We are, therefore, forced to regard the
constitution of the wild _grossulariata_ female as identical with that of
the F_1 female, _i.e._ as heterozygous for the _grossulariata_ factor as
well as for the factor for femaleness. Though from a region where
_lacticolor_ is unknown, the "pure" wild _grossulariata_ female is
nevertheless a permanent mongrel, but it can never reveal its true colours
unless it is mated with a male which is either heterozygous for G or pure
_lacticolor_. And as all the wild northern males are {105} pure for the
_grossulariata_ character this can never happen in a state of nature.
[Illustration: FIG. 19.
Scheme illustrating the result of crossing a Silky hen with a Brown Leghorn
cock. Black sex signs denote deeply pigmented birds, and light sex signs
those without pigmentation. The light signs with a black dot in the centre
denote birds with a small amount of pigment.]
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