Mendelism: Third EditionPunnett, Reginald Crundall
Science
Mendelism: Third Edition
Punnett, Reginald Crundall
Mendel's law
though such work was almost entirely confined to the botanists. Apart from
the fact that plants lend themselves to hybridisation work more readily
than animals, there was probably another reason why zoologists neglected
this form of investigation. The field of zoology is a wider one than that
of botany, presenting a far greater variety of type and structure. Partly
owing to their importance in the study of medicine, and partly owing to
their smaller numbers, the anatomy of the vegetable was far better known
than that of the animal kingdom. It is, therefore, not surprising that the
earlier part of the nineteenth century found the zoologists, under the
influence of Cuvier and his pupils, devoting their entire energies to
describing the anatomy of the new forms of animal life which careful search
at home and fresh voyages of discovery abroad were continually bringing to
light. During this period the zoologist had little inclination or
inducement to carry on those investigations in hybridisation which were
occupying the attention of some botanists. Nor did the efforts of the
botanists afford much {10} encouragement to such work, for in spite of the
labour devoted to these experiments, the results offered but a confused
tangle of facts, contributing in no apparent way to the solution of the
problem for which they had been undertaken. After half a century of
experimental hybridisation the determination of the relation of species and
varieties to one another seemed as remote as ever. Then in 1859 came the
_Origin of Species_, in which Darwin presented to the world a consistent
theory to account for the manner in which one species might have arisen
from another by a process of gradual evolution. Briefly put, that theory
was as follows: In any species of plant or animal the reproductive capacity
tends to outrun the available food supply, and the resulting competition
leads to an inevitable struggle for existence. Of all the individuals born,
only a portion, and that often a very small one, can survive to produce
offspring. According to Darwin's theory, the nature of the surviving
portion is not determined by chance alone. No two individuals of a species
are precisely alike, and among the variations that occur some enable their
possessors to cope more successfully with the competitive conditions under
which they exist. In comparison with their less favoured brethren they have
a better chance of surviving in the struggle for existence and consequently
of leaving offspring. The argument is completed by the further assumption
of a principle of heredity, in virtue of which offspring tend to {11}
resemble their parents more than other members of the species. Parents
possessing a favourable variation tend to transmit that variation to their
offspring, to some in greater, to others in less degree. Those possessing
it in greater degree will again have a better chance of survival, and will
transmit the favourable variation in even greater degree to some of their
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