Although the Euploeinae are not represented on the African continent, it is
the headquarters of another distasteful family of butterflies--the
Acraeinae--which is but sparingly represented in the Oriental region[17].
{34} Of smaller size than the Danaines they are characterised, like this
group, by their tenacity of life and by the presumably distasteful
character of their body juices. They are said also to possess an offensive
odour apparently exuded through the thorax. The majority of the members of
the group fall into the two genera _Acraea_ and _Planema_. Species of
Acraea are on the whole characterised by their general bright red-brown
colour and by the conspicuous black spots on both fore and hind wings. A
typical Acraeine pattern is that of _Acraea egina_ (Pl. VI, fig. 7) which
is mimicked remarkably closely by the Nymphaline _Pseudacraea boisduvali_
and by the Swallow-tail _Papilio ridleyanus_ (Pl. VI, figs. 5 and 6).
In the genus _Planema_ the spots are as a rule fewer and clustered near the
body, while on both fore and hind wings there is a tendency to develop
clear wide band-like areas of orange or white (cf. Pl. VII).
Like the Acraeas the Planemas are principally mimicked by species of
_Pseudacraea_ and of _Papilio_. Some of the cases of resemblance between
_Planema_ and _Pseudacraea_ are among the most striking known. _Planema
macarista_ is one of those comparatively rare instances in which a model
shews a marked difference in the pattern of the two sexes. The clear area
on the fore wing of the male is deep orange, whereas in the female it is
somewhat different in shape, and, like the area on the hind wing, is white
(cf. Pl. VII, figs. 1 and 2). {35} _Pseudacraea eurytus hobleyi_ (Pl. VII,
figs. 6 and 7) shews a similar difference in the sexes, the male and female
of this species mimicking respectively the male and female of _Planema
macarista_. The case is made even more remarkable by the fact that both of
the sexual forms of _Planema macarista_ are mimicked by the Satyrine
_Elymnias phegea_ (Pl. VII, fig. 9), though in this species either the
black and white, or the black, white, and orange form may occur in either
sex. Among the best Papilionine mimics of the Planemas is _Papilio cynorta_
whose female is extraordinarily like the common _Planema epaea_ (Pl. VII,
figs. 5 and 10). The resemblance of the _planemoides_ female of _P.
dardanus_ to _P. poggei_ has already been noticed.
A striking feature of the African continent is the frequency with which
mimetic forms are found among the Lycaenidae. As a rule the "blues" rarely
exhibit mimetic analogies, but in Africa there are several species,
especially those of the genus _Mimacraea_, which closely resemble
Acraeines. Others again bear a marked resemblance to certain small
Pierines, _Citronophila similis_ from S. Nigeria for example being
extraordinarily like the common _Terias brigitta_, a small bright yellow
Pierine with black-edged wings.
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