(1) That the imitative species occur in the same area and occupy the very
same station as the imitated.
(2) That the imitators are always the more defenceless. {51}
(3) That the imitators are always less numerous in individuals.
(4) That the imitators differ from the bulk of their allies.
(5) That the imitation, however minute, is _external_ and _visible_ only,
never extending to internal characters or to such as do not affect the
external appearance.
In offering certain criticisms of the mimicry explanation it will be
convenient to do so in connection with these five conditions which Wallace
regarded as constant for all cases of mimetic resemblance.
(1) _That the imitative species occur in the same area and occupy the very
same station as the imitated._
This on the whole is generally true. It is well shewn in some of the most
striking cases such as those of the Old-World Papilios that mimic Danaines,
or of the Dismorphias and their Ithomiine models. In many of these cases
the range of neither model nor mimic is a very wide one, yet the mimic is
found strictly inside the area inhabited by the model. _Papilio agestor_,
for instance, is only found where _Caduga tytia_ occurs, nor is _P. mendax_
known outside the area frequented by _Euploea rhadamanthus_. Even more
striking in this respect are some of the Ithomiine-Dismorphia resemblances
in the New World. The Ithomiine models are as a rule very local though very
abundant. Two hundred miles away the predominant Ithomiine often bears
quite a distinct pattern, and when this is the case the mimicking
_Dismorphia_ is generally changed in the same sense. {52} But though mimic
and model may be found together in the same locality, they do not always
occupy the same station in the sense that they fly together. According to
Seitz[25] the Dismorphias themselves do not fly with the Ithomiines which
they mimic. The occurrence of butterflies is largely conditioned by the
occurrence of the plants on which the larva feeds, and this is especially
true of the female, which, as has already been noticed, is more commonly
mimetic than the male. The female of _Papilio polytes_, for instance, is
found flying where are to be found the wild citronaceous plants on which
its larva feeds. On the other hand, its so-called models, _Papilio hector_
and _P. aristolochiae_, are generally in the proximity of the Aristolochias
on which their larvae feed. The two plants are not always found together,
so that one frequently comes across areas where _P. polytes_ is very
abundant while the models are scarce or absent.
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