There are writers on these questions of mimicry who adopt a view more or
less intermediate between those just discussed. They regard the resemblance
as having arisen in the first place as a sport of some magnitude on the
part of the mimic, rendering it sufficiently like the model to cause some
confusion between the two. A rough-hewn resemblance is first brought about
by a process of mutation. Natural selection is in this way given something
to work on, and forthwith proceeds to polish up the resemblance until it
becomes exceedingly close. Natural selection does not originate the
likeness, but, as soon as a rough one has made its appearance, it comes
into operation and works it up through intermediate stages into the
finished portrait. It still plays some part in the formation of a mimetic
resemblance though its rôle is now restricted to the putting on of the
finishing touches. Those who take this view hold also that the continued
action of natural selection is necessary in order to keep the likeness up
to the mark. They suppose that if selection ceases the likeness gradually
deteriorates owing to the coming into operation of a mysterious {72}
process called regression. This idea involves certain conceptions as to the
nature of variation which we shall discuss later.
Though it is difficult to regard Batesian mimicry as produced by the
accumulation of small variations through natural selection, it is perhaps
rather more plausible to suppose that such a process may happen in
connection with the numerous instances of Müllerian mimicry. For since the
end result is theoretically to the advantage of both species instead of but
one, it is possible to argue that the process would be simplified by their
meeting one another halfway, as Müller[34] himself originally suggested.
Variations on the part of each in the direction of the other would be
favourably selected, the mimicry being reciprocal.
Difficulties, however, begin to arise when we inquire into the way in which
this unification of pattern may be conceived of as having come about. By no
one have these difficulties been more forcibly presented than by
Marshall[35] in an able paper published a few years ago, and perhaps the
best way of appreciating them is to take a hypothetical case used by him as
an illustration.
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