In Europe, in spite of the fact that the winter climate is much milder than in corresponding latitudes in North America, the number of kinds of forest trees is much less, there is little resemblance to those of Asia and the eastern United States, and the trees have generally a less southern aspect. This is the more remarkable in that trees of southern facies introduced from China and Japan and from the United States thrive admirably in Europe, showing that there is no climatic obstacle to their presence there. To mention only a few examples, the Tree of Heaven (_Ailanthus glandulosa_), so very common, even as a wild tree in many parts of the continent of Europe, was introduced from China, while the beautiful _Sophora japonica_, so frequently planted in towns, comes, as its name indicates, from Japan, and the various species of those beautiful flowering trees known as Catalpa are either American or Asiatic. The western plane (_Platanus occidentalis_), another favourite town tree, comes from the United States, and other American trees which are found very abundantly in towns in the warmer parts of Europe are the black walnut and the honey locust (_Gleditschia tricanthos_). Perhaps more striking than any of these is the case of the so-called false acacia (_Robinia pseudacacia_), which is as common over a great part of the continent of Europe as hawthorn bushes or wild roses are with us, and yet is a North American species, introduced less than three hundred years ago. Generally, we may say that all the more beautiful trees now growing in the warmer parts of Europe come either from eastern Asia or from the United States. In other words, the Ice Age seems to have greatly impoverished the flora of Europe. To a less extent this is also true of western North America, which has fewer species of trees than the east. Why had the ice this impoverishing effect upon Europe? The topography of the continent supplies the answer. In the first place, in Europe there are numerous transverse chains of mountains. The Pyrenees, the Alps, the Caucasus, each with its load of ice, each with glaciers deploying on the low ground at its feet, must have been obstacles in the way of the southern migration alike of plants and of animals. Again, even if these obstacles were passed or turned, the great inland sea formed another barrier further south. In consequence of this difficulty in finding asylums the pre-glacial plants and animals must have perished in considerable numbers, and thus a general impoverishment took place. One must not of course exaggerate. A proportion of the pre-glacial forms did succeed in living through the period of stress, but many must have been, as it were, squeezed out of Europe or out of existence by the unfavourable climatic conditions.