What plant has the most complex single stigma and pistil? The most
complex I, in my ignorance, can think of is in Iris. I want to know
whether anything beats in modification the rostellum of Catasetum.
To-morrow I mean to be at Catasetum. Hurrah! What species is it? It
is wonderfully different from that which Veitch sent me, which was C.
saccatum.
According to the vessels, an orchid flower consists of three sepals and
two petals free; and of a compound organ (its labellum), consisting
of one petal and of two (or three) modified anthers; and of a second
compound body consisting of three pistils, one normal anther, and two
modified anthers often forming the sides of the clinandrum.
LETTER 604. TO JOHN LINDLEY.
(604/1. It was in the autumn of 1861 that Darwin made up his mind to
publish his Orchid work as a book, rather than as a paper in the Linnean
Society's "Journal." (604/2. See "Life and Letters," III., page 266.)
The following letter shows that the new arrangement served as an
incitement to fresh work.)
Down, October 25th [1861?]
Mr. James Veitch has been most generous. I did not know that you had
spoken to him. If you see him pray say I am truly grateful; I dare not
write to a live Bishop or a Lady, but if I knew the address of "Rucker"?
and might use your name as introduction, I might write. I am half mad on
the subject. Hooker has sent me many exotics, but I stopped him, for I
thought I should make a fool of myself; but since I have determined to
publish I much regret it.
(FIGURE 9.--HABENARIA CHLORANTHA (Longitudinal course of bundles).)
(605/1. The three upper curved outlines, two of which passing through
the words "upper sepal," "upper petal," "lower sepal," were in red in
the original; for explanation see text.)
LETTER 605. TO J.D. HOOKER.
(605/2. The following letter is of interest because it relates to one of
the two chief difficulties Darwin met with in working out the morphology
of the orchid flower. In the orchid book (605/3. Edition I., page 303.)
he wrote, "This anomaly [in Habenaria] is so far of importance, as it
throws some doubt on the view which I have taken of the labellum being
always an organ compounded of one petal and two petaloid stamens." That
is to say, it leaves it open for a critic to assert that the vessels
which enter the sides of the labellum are lateral vessels of the petal
and do not necessarily represent petaloid stamens. In the sequel he
gives a satisfactory answer to the supposed objector.)
Down, November 10th, [1861].
Public-domain text, read in full here on John Shaqi.
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