For the love of God help me. I believe all my work (about a
fortnight) is useless. Look at this accursed diagram (Figure 9) of the
butterfly-orchis [Habenaria], which I examined after writing to you
yesterday, when I thought all my work done. Some of the ducts of the
upper sepal (605/4. These would be described by modern morphologists as
lower, not upper, sepals, etc. Darwin was aware that he used these terms
incorrectly.) and upper petal run to the wrong bundles on the column. I
have seen no such case.
This case apparently shows that not the least reliance can be placed on
the course of ducts. I am sure of my facts.
There is great adhesion and extreme displacement of parts where the
organs spring from the top of the ovarium. Asa Gray says ducts are very
early developed, and it seems to me wonderful that they should pursue
this course. It may be said that the lateral ducts in the labellum
running into the antero-lateral ovarian bundle is no argument that the
labellum consists of three organs blended together.
In desperation (and from the curious way the base of upper petals are
soldered at basal edges) I fancied the real form of upper sepal, upper
petal and lower sepal might be as represented by red lines, and that
there had been an incredible amount of splitting of sepals and petals
and subsequent fusion.
This seems a monstrous notion, but I have just looked at Bauer's drawing
of allied Bonatea, and there is a degree of lobing of petals and sepals
which would account for anything.
Now could you spare me a dry flower out of your Herbarium of Bonatea
speciosa (605/5. See "Fertilisation of Orchids," Edition I., page 304
(note), where the resemblances between the anomalous vessels of Bonatea
and Habenaria are described. On November 14th, 1861, he wrote to Sir
Joseph: "You are a true friend in need. I can hardly bear to let the
Bonatea soak long enough."), that I might soak and look for ducts. If
I cannot explain the case of Habenaria all my work is smashed. I was a
fool ever to touch orchids.
LETTER 606. TO J.D. HOOKER. Down, November 17th [1861].
Public-domain text, read in full here on John Shaqi.
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