Produced by Chris Curnow, Tom Cosmas, Joseph Cooper and the Online Distributed Proofreading Team at http://www.pgdp.net UNIVERSITY OF KANSAS PUBLICATIONS MUSEUM OF NATURAL HISTORY Volume 11, No. 10, pp. 527-669, 16 pls., 29 figs. March 7, 1960 Natural History of the Ornate Box Turtle, Terrapene ornata ornata Agassiz BY JOHN M. LEGLER UNIVERSITY OF KANSAS LAWRENCE 1960 UNIVERSITY OF KANSAS PUBLICATIONS, MUSEUM OF NATURAL HISTORY Editors: E. Raymond Hall, Chairman, Henry S. Fitch, Robert W. Wilson Volume 11, No. 10, pp. 527-669, 16 pls., 29 figs. Published March 7, 1960 UNIVERSITY OF KANSAS Lawrence, Kansas PRINTED IN THE STATE PRINTING PLANT TOPEKA, KANSAS 1960 [Union Label] 28-773 Natural History of the Ornate Box Turtle, Terrapene ornata ornata Agassiz BY JOHN M. LEGLER CONTENTS PAGE Introduction 531 Acknowledgments 531 Systematic Relationships and Distribution 532 Fossils 534 Economic Importance 534 Study Areas 535 Materials and Methods 537 Terminology 539 Habitat and Limiting Factors 539 Habitat in Kansas 542 Reproduction 543 Mating 543 Insemination 545 Sexual Cycle of Males 545 Sexual Cycle of Females 549 Nesting 554 Eggs 558 Embryonic Development 560 Fertility and Prenatal Mortality 564 Reproductive Potential 565 Number of Reproductive Years 565