Natural History of the Racer Coluber constrictorFitch, Henry S. (Henry Sheldon)
Science
Natural History of the Racer Coluber constrictor
Fitch, Henry S. (Henry Sheldon)
Racer snake
Kansas, and from Cedar Bluff Reservoir, 23 miles west and seven miles
south of Hays in Trego County, western Kansas, where live-trapping was
carried on in 1959 and 1960. Additional data were obtained on
numerous field trips to various collecting localities in northeastern
Kansas. My first-hand knowledge of the species is also based, in part,
on many years of field experience with the far western subspecies _C.
c. mormon_ in western Oregon and California, and on similar experience
in 1947 and 1948 with the southern subspecies, _C. c. anthicus_ in
central Louisiana.
This varied field experience with the species at localities well
scattered throughout its geographic range has added perspective to the
study even though most of the records were collected within a radius
of three quarters of a mile. No one locality can be regarded as
entirely typical of a species' habitat over its range as a whole.
According to my philosophy, the ecological niche of a species is
subject to geographical variation analogous to the variation to be
seen in the morphological characters of the animal itself. Different
community associates, including different competitors, prey, and
predators, and different physical factors enforce a somewhat different
way of life on a species in geographically remote parts of its range.
When analyzed these differences often turn out to have a genetic
basis. Thus, limits of tolerance to heat, cold, and drought often vary
geographically, and the population density, reproductive potential,
seasonal cycle, and other properties of populations may be altered
either by the direct effect of the environment, or through its effect
on the genetic constitution, produced by natural selection.
The local population of racers studied was near the center of the
species' geographic range, and is to some extent representative of the
species as a whole, though differing in its ecology from other
populations in proportion to their remoteness and the distinctness of
their habitats. It has not been demonstrated that ecological traits of
populations change in a discontinuous manner or correspond in their
limits with those of named subspecies. More likely geographical
variation is continuous and parallels morphological variation only in
a general way. Certainly the boundaries of subspecies' ranges should
not be accorded undue emphasis in an ecological study.
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