ANALYSIS OF MORPHOLOGICAL CHARACTERS 330 Osteology 330 _Descriptive Osteology of Smilisca baudini_ 331 _Developmental Cranial Osteology of Smilisca baudini_ 333 _Comparative Osteology_ 336 Musculature 341 Skin 342 _Structure_ 342 _Comparative Biochemistry of Proteins_ 343 External Morphological Characters 343 _Size and Proportions_ 343 _Shape of Snout_ 344 _Hands and Feet_ 344 _Ontogenetic Changes_ 344 Coloration 344 _Metachrosis_ 345 Chromosomes 345 NATURAL HISTORY 345 Breeding 345 _Time of Breeding_ 345 _Breeding Sites_ 346 _Breeding Behavior_ 346 _Breeding Call_ 351 Eggs 356 Tadpoles 357 _General Structure_ 357 _Comparison of Species_ 357 _Growth and Development_ 361 _Behavior_ 365 PHYLOGENETIC RELATIONSHIPS 366 Interspecific Relationships 366 Evolutionary History 369 SUMMARY AND CONCLUSIONS 371 LITERATURE CITED 372 INTRODUCTION