Neotropical Hylid Frogs, Genus SmiliscaDuellman, William Edward
Science
Neotropical Hylid Frogs, Genus Smilisca
Duellman, William Edward
Amphibians -- Central America; Frogs; Smilisca
_Diagnosis._--Size small ([M] 38.0 mm., [F] 46.0 mm.), differing from
other species in the genus by the following combination of characters:
skull about as long as broad; frontoparietal fontanelle keyhole-shaped;
supraorbital flanges absent; squamosal small, not in contact with
maxillary; bony portion of ethmoid terminating at anterior edge of
orbit; tarsal fold weak, two-thirds length of tarsus; inner metatarsal
tubercle small, low, flat, elliptical; snout rounded in dorsal profile;
lips thin and flaring; fingers having only vestige of web; toes one-half
webbed; diameter of tympanum about two-thirds that of eye; narrow labial
stripe white; pair of dark brown (sometimes interconnected) stripes on
tan dorsum; no blue spots on flanks or thighs; vocal sac in breeding
males pale brown. (Foregoing combination of characters distinguishing
_S. puma_ from other species in genus.)
_Description and variation._--Ten breeding males from the vicinity of
Puerto Viejo, Heredia Province, Costa Rica, have snout-vent lengths of
32.5 to 37.9 mm. (34.8 mm.). In these specimens, the length of the tibia
to the snout-vent length is 0.48 to 0.53 (0.51), and the tympanum/eye
ratio is 0.52 to 0.72 (0.65). Seven females have snout-vent lengths of
40.8 to 45.8 mm. (43.9 mm.). No individual has more than a vestige of a
web between the second and third and fourth fingers. None has a web
between the first and second fingers. Breeding males lack nuptial
excrescences on the thumbs. The vocal sac is moderately large and
bilobed.
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