Neotropical Hylid Frogs, Genus SmiliscaDuellman, William Edward
Science
Neotropical Hylid Frogs, Genus Smilisca
Duellman, William Edward
Amphibians -- Central America; Frogs; Smilisca
_Diagnosis._--Size moderate ([M] 45.0 mm., [F] 62.2 mm.); skull wider
than long, having large, ovoid frontoparietal fontanelle; supraorbital
flanges absent; squamosal small, not contacting maxillary; bony section
of ethmoid extending anteriorly between nasals; tarsal fold weak, full
length of tarsus; inner metatarsal tubercle low, flat, elliptical; lips
thick, rounded, not flaring; fingers one-third webbed; toes
three-fourths webbed; diameter of tympanum about one-half that of eye;
margin of upper lip faintly marked by interrupted creamy white stripe;
dark spots on dorsum; pale flecks on flanks and posterior surfaces of
thighs; vocal sacs in breeding males dark brown. (Foregoing combination
of characters distinguishing _S. sila_ from any other species in genus.)
_Description of holotype._--Snout-vent length 36.6 mm.; tibia length
19.8 mm., 54.1 per cent of snout-vent length; foot length 15.5 mm., 42.3
per cent of snout-vent length; head length 12.7 mm., 34.7 per cent of
snout-vent length; head width 13.3 mm., 36.8 per cent of snout-vent
length; snout short, in lateral profile truncate, only slightly rounded
above, in dorsal profile rounded; canthus rounded; loreal region
noticeably concave; lips thick, rounded, not flaring; nostrils not
protuberant, directed laterally; internarial distance 3.0 mm.;
internarial area flat; top of head flat; interorbital distance 3.5 mm.,
26.3 per cent of head width; diameter of eye 4.2 mm., thrice distance
(1.4 mm.) from tympanum to eye, and half again distance (2.8 mm.) from
orbit to nostril; pupil horizontally ovoid; width of eyelid 2.8 mm.,
21.1 per cent of head width; dermal fold from posterior corner of orbit
covering upper edge of tympanum to point above insertion of forelimb;
diameter of tympanum 2.3 mm., 54.7 per cent of diameter of eye; no
axillary membrane; arms moderately robust; weak fold on wrist; faintly
scalloped fold along ventrolateral margin of forearm; fingers short,
slender; fingers from shortest to longest, 1-2-4-3; vestige of web
between first and second fingers; others about two-fifths webbed; discs
moderate, diameter of that on third finger about one-third diameter of
eye; triangular outer palmar tubercle; elliptical inner palmar tubercle
on base of pollex; subarticular tubercles large, conical, none bifid;
supernumerary tubercles few, large, conical; brown nuptial excrescence
on prepollex; heels overlap by about one-fifth length of shank when hind
limbs adpressed; tibiotarsal articulation extending to nostril; tarsal
fold weak, extending nearly full length of tarsus; inner metatarsal
tubercle elliptical, flat; outer metatarsal tubercle absent; toes
moderately long; toes from shortest to longest, 1-2-3-5-4, third and
fifth about equal in length; discs about same size as those on fingers;
webbing extending to middle of penultimate phalanx on all toes, except
only to distal end of antepenultimate phalanx of fourth toe;
subarticular tubercles round; supernumerary tubercles large, round,
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