Neotropical Hylid Frogs, Genus SmiliscaDuellman, William Edward
Science
Neotropical Hylid Frogs, Genus Smilisca
Duellman, William Edward
Amphibians -- Central America; Frogs; Smilisca
_Diagnosis._--Size moderate ([M] 45 mm., [F] 64 mm.); skull slightly
wider than long, having large and elongate frontoparietal fontanelle;
supraorbital flanges absent; squamosal small, not contacting maxillary;
bony section of ethmoid terminating just anterior to anterior edge of
orbit; tarsal fold weak, full length of tarsus; inner metatarsal
tubercle long, low, flat, elliptical; lips thin and flaring; fingers
one-half webbed; toes four-fifths webbed; diameter of tympanum about
one-half that of eye; no white labial stripe; dorsal dark markings
irregular, sometimes forming broad transverse bars; pale flecks on
flanks and usually on posterior surfaces of thighs; vocal sacs in
breeding males white. (Foregoing combination of characters
distinguishing _S. sordida_ from any other species in genus.)
_Description and variation._--Ten breeding males from 15 to 20
kilometers west-southwest of San Isidro el General, San Jose, Costa
Rica, have snout-vent lengths of 38.1 to 42.6 mm. (40.5 mm.). In these
specimens, the tibia/snout-vent length ratio is 0.50 to 0.54 (0.52), and
the tympanum/eye ratio is 0.45 to 0.57 (0.49). Specimens from the
Pacific slopes of Costa Rica are larger than those from the Meseta
Central and the Caribbean lowlands. Ten males from 6 kilometers east of
Golfito, Puntarenas, have snout-vent lengths of 38.4 to 44.6 mm. (41.8
mm.), and five males from Rincon, Peninsula de Osa, have snout-vent
lengths of 38.8 to 41.6 mm. (40.3 mm.). Snout-vent lengths of ten males
from La Fortuna, Alajuela, are 31.9 to 36.0 mm. (34.4 mm.), of ten males
from Pandora, Limon, 33.8 to 37.6 mm. (35.9 mm.), and of ten males from
Escazu and Rio Jorco on the Meseta Central, 34.3 to 37.6 mm. (36.0 mm.).
Eight females from the Rio Jorco on the Meseta Central have snout-vent
lengths of 48.8 to 53.8 mm. (50.4 mm.), and six females from various
localities on the Pacific slopes of Costa Rica have snout-vent lengths
of 56.5 to 64.0 mm. (59.8 mm.). The only noticeable differences in
proportions between males and females is in the tympanum/eye ratio; for
example, this ratio is 0.47 to 0.53 (0.49) and 0.54 to 0.68 (0.61) in
ten males and eight females, respectively, from the Meseta Central.
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