North American Recent Soft-Shelled Turtles (Family Trionychidae) — John Shaqi
North American Recent Soft-Shelled Turtles (Family Trionychidae)Webb, Robert G. (Robert Gravem)
Science
North American Recent Soft-Shelled Turtles (Family Trionychidae)
Webb, Robert G. (Robert Gravem)
Soft-shelled turtles
Large females of _T. spinifer_ have obtuse prominences in the center
of the carapace posteriorly, some of which in many specimens are
arranged in longitudinal rows; I cannot discern any correlation of
number or arrangement of prominences with size in _spinifer_ or
_ferox_. The carapace in adult males of _spinifer_ bears small, sharp
tubercles that make the surface feel like sandpaper. The tubercles on
the anterior edge of the carapace in adults of both sexes vary from
round to equilateral and conical to low and flattened (see comments on
tuberculation under subsection entitled "Geographic Variation"). Some
large females of the same subspecies have tubercles on the anterior
edge of the carapace that may be conical (higher than wide) or
equilateral. The difference in shape of the tubercles seems not to be
correlated with size because one _T. s. pallidus_, 30.5 centimeters
(TU 13212) has prominent but blunted and equilateral tubercles,
whereas, another female of _pallidus_, 20.8 centimeters (TU 13210),
from the same locality has higher, conical tubercles. The blunted,
equilateral tubercles may be the result of environmental wear, or the
difference in shape of tubercles may be due to individual variation.
_Pattern on Carapace_
Two features of the pattern on the carapace are of taxonomic worth: 1)
the width and distinctness of the pale rim at the periphery of the
carapace (marginal rim), if present, and 2) the kind of pattern on the
carapace (juvenal pattern). The marginal rim is absent in females of
_T. ater_, and only faintly evident in males. The marginal rim is
obscured or absent (adult males and females) and is not separated from
the ground color of the carapace by a dark marginal line in hatchlings
of _T. ferox_. The carapace of _T. muticus_ has a marginal rim that is
usually separated from the ground color of the carapace by an
ill-defined, dark marginal line; some individuals lack the marginal
dark line. The subspecies of _T. spinifer_ have a well-defined, dark,
marginal line that separates the marginal rim from the ground color of
the carapace; _T. s. asper_ has more than one dark marginal line on
the carapace. The marginal rim is ill-defined and blotched, or absent,
in large females of all species of _Trionyx_.
The marginal rim is widest at the posterior end of the carapace and
lacking in the nuchal area. The width of the pale marginal rim is very
narrow, almost to the degree of being absent, in juveniles of _T.
ferox_. _T. s. emoryi_ has a pale, marginal rim that is four or five
times wider posteriorly than it is laterally, whereas posteriorly the
width of the rim in the other subspecies of _T. spinifer_ and in the
species _T. muticus_ is only two or three times wider posteriorly than
it is laterally.
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