Strata of limestone, 2,500 feet in thickness, were accumulated in the ocean by the labours of Coral-polypes, allied to, but totally distinct from, those which had previously existed in the primary system. On the floor of a shallow sea, which then occupied the middle of what is now England, the coral reefs rose perpetually towards the day, atom by atom, the strata on which they were founded slowly and steadily sinking ever to a lower level, while successive generations of the industrious zoophytes wrought upwards, to maintain their position within reach of the light and warmth. What period of time was requisite for the aggregation of coral structure to the perpendicular thickness of 2,500 feet? While this was going on, other Invertebrata were living in the shallow seas, mostly differing from the older species, which had become by this time extinct. Encrinites and Sea-urchins existed; some _Foraminifera_ were astonishingly abundant; the _Cephalopoda_ and the _Brachiopoda_ presented a vast variety of species; and about seventy sorts of Fishes, mostly Sharks, characterised the age. On the coral limestone lies a sort of conglomerate, known as the millstone grit; and on this is laid that source of Britain's eminence, the _coal_. The coal measures of South Wales are estimated at 12,000 feet in thickness. The profusion of vegetable life that must have combined to make the coal in these, has no parallel in this age; no, not in the teeming forests of South America, or the great isles of the Oriental Archipelago. The circumstances which favoured this enormous development of plants, seem never to have been repeated in subsequent ages, since the coal measures which are found in the later strata are thin and inconsiderable, compared with those we are considering. M. Adolphe Brogniart suggests that in this period, from some source or other, carbonic acid was generated in vast abundance; or, at least, that it existed in the air, in a far greater proportion than it does now; and it is singularly confirmatory of his view, that terrestrial animals, to which this gas is fatal, have left almost no traces of their existence, during the age of these vast forests--a circumstance otherwise strange and unaccountable. "Those parts," says Mr. Ansted, "of the great carboniferous series which generally include the beds of coral, consist of muddy and sandy beds, alternating with one another, and with the coal itself. Some of them would appear to be of fresh-water, and some of marine origin; and they abound, for the most part, with remains of the leaves of Ferns and fern-like trees, together with the crushed trunks of these and other trees, whose substance may have contributed to form the great accumulations of bituminised and other vegetable carbon obtained from these strata.