On Germinal Selection as a Source of Definite Variation — John Shaqi
On Germinal Selection as a Source of Definite VariationWeismann, August
Science
On Germinal Selection as a Source of Definite Variation
Weismann, August
Variation (Biology)
The facts do indeed exist and lie before our very eyes, despite their not
having been recognised as such before. All _artificial selection_ practised
by man rests on the fact that by means of the selection of individuals
having a given character slightly more pronounced than usual, there is
gradually produced a general augmentation of this character, which
subsequently reaches a point never before attained by any individual {34}
of this species. I shall choose an example which seems to me especially
clear and simple because only one character has been substantially modified
here. The long-tailed variety of domestic cock, now bred in Japan and
Corea, owes its existence to skilful selection and not at all to the
circumstance that at some period of the race's history a cock with
tail-feathers six feet in length suddenly and spasmodically appeared. At
the present day even, as Professor Ishikawa of Tokio writes me, the
breeders still make extraordinary efforts to increase the length of the
tail, and every inch gained adds considerably to the value of the bird. Now
nothing has been done here whatever except always to select for purposes of
breeding the cocks with the longest feathers; and in this way alone were
these feathers, after the lapse of many generations, prolonged to a length
far exceeding every previous variation.
I once asked a famous dove-fancier, Mr. W. B. Tegetmeier of London, whether
it was his opinion that by artificial selection alone a character could be
augmented. He thought a long time and finally said: "It is without our
power to do anything if the variation which we seek is not presented, but
once that variation is given, then I think the augmentation can be
effected." And that in fact is the case. If cocks had never existed whose
tail-feathers were a little longer than usual the Japanese breed could
never have originated; but as the facts are, always the cocks with the
longest feathers were chosen from each generation, and these only were
bred, and thus a hereditary augmentation of the character in question was
effected, which would hardly have been deemed possible.
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