On Germinal Selection as a Source of Definite VariationWeismann, August
Science
On Germinal Selection as a Source of Definite Variation
Weismann, August
Variation (Biology)
In this way we acquire an approximate conception of the possible mechanical
_modus operandi_ of actual events--namely, of the manner in which the
useful variations required by the conditions of life _can_ always, that is,
very frequently, make their appearance. This possibility is the sole
condition of our being able to understand how different parts of the body,
absolutely undefined in extent, can appear as variational units and vary in
the same or in different directions, according to the special needs of the
case, or as the conditions of life prescribe. Thus, for example, in the
case of the butterfly's wings it rests entirely with utility to decide the
size and the shape of the spots that shall vary simultaneously in the same
direction. At one time the whole under surface of the wing appears as the
variational unit and has the same color; at another the inside half, which
is dark, is contrasted with the outside half which is bright; or the same
contrast will exist between the anterior and posterior halves; or, finally,
narrow stripes or line-shaped streaks will behave as variational units and
form contrasts with manifold kinds of spots or with the broader intervals
between them, with the result that the picture of a leaf or of another
protected species is produced.
I must refrain from entering into the details of such cases and shall
illustrate my views regarding the color-transformations of butterflies'
wings by the simplest {49} conceivable example--viz. that of the uniform
change of color on the entire under surface of the wing.
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