This portion of matter, then, is kept together by the intermolecular
force of cohesion; in the movements of its particles relatively to
one another, and in its own movements relative to adjacent matter, it
meets with the opposing force of friction. It is acted on by gravity,
and this force tends (though slightly, owing to the Amoeba’s small
mass, and to the small difference between its density and that of the
surrounding fluid), to flatten it down upon the solid substance on
which it may be creeping. Our Amoeba tends, in the next place, to be
deformed by any pressure from outside, even though slight, which may be
applied to it, and this circumstance shews it to consist of matter in a
fluid, or at least semi-fluid, state: which state is further indicated
when we observe streaming or current motions in its interior. {13}
Like other fluid bodies, its surface, whatsoever other substance, gas,
liquid or solid, it be in contact with, and in varying degree according
to the nature of that adjacent substance, is the seat of molecular
force exhibiting itself as a surface-tension, from the action of which
many important consequences follow, which greatly affect the form of
the fluid surface.
While the protoplasm of the Amoeba reacts to the slightest pressure,
and tends to “flow,” and while we therefore speak of it as a fluid,
it is evidently far less mobile than such a fluid, for instance, as
water, but is rather like treacle in its slow creeping movements as
it changes its shape in response to force. Such fluids are said to
have a high viscosity, and this viscosity obviously acts in the way of
retarding change of form, or in other words of retarding the effects
of any disturbing action of force. When the viscous fluid is capable
of being drawn out into fine threads, a property in which we know that
the material of some Amoebae differs greatly from that of others, we
say that the fluid is also _viscid_, or exhibits viscidity. Again, not
by virtue of our Amoeba being liquid, but at the same time in vastly
greater measure than if it were a solid (though far less rapidly than
if it were a gas), a process of molecular diffusion is constantly going
on within its substance, by which its particles interchange their
places within the mass, while surrounding fluids, gases and solids in
solution diffuse into and out of it. In so far as the outer wall of
the cell is different in character from the interior, whether it be
a mere pellicle as in Amoeba or a firm cell-wall as in Protococcus,
the diffusion which takes place _through_ this wall is sometimes
distinguished under the term _osmosis_.
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