When we have four bubbles in conjunction, they would seem to be capable
of arrangement in two symmetrical ways: either, as in Fig. 116 (A),
with the four partition-walls meeting at right angles, or, as in (B),
with _five_ partitions meeting, three and three, at angles of 120°.
This latter arrangement is strictly analogous to the arrangement of
three bubbles in Fig. 114. Now, though both of these figures, from
their symmetry, are apparently figures of equilibrium, yet, physically,
the former turns out to be of unstable and the latter of stable
equilibrium. If we try to bring our four bubbles into the form of Fig.
116, A, such an arrangement endures only for an instant; the partitions
glide upon each other, a median wall springs into existence, and the
system at once assumes the form of our second figure (B). This is a
direct consequence of the law of minimal areas: for it can be shewn, by
somewhat difficult mathematics (as was first done by Lamarle), that,
in dividing a closed space into a given number of chambers by means
of partition-walls, the least possible area of these partition-walls,
taken together, can only be attained when they meet together in groups
of three, at equal angles, that is to say at angles of 120°. {310}
Wherever we have a true cellular complex, an arrangement of cells in
actual physical contact by means of a boundary film, we find this
general principle in force; we must only bear in mind that, for its
perfect recognition, we must be able to view the object in a plane
at right angles to the boundary walls. For instance, in any ordinary
section of a vegetable parenchyma, we recognise the appearance of
a “froth,” precisely resembling that which we can construct by
imprisoning a mass of soap-bubbles in a narrow vessel with flat sides
of glass; in both cases we see the cell-walls everywhere meeting, by
threes, at angles of 120°, irrespective of the size of the individual
cells: whose relative size, on the other hand, determines the
_curvature_ of the partition-walls. On the surface of a honey-comb we
have precisely the same conjunction, between cell and cell, of three
boundary walls, meeting at 120°. In embryology, when we examine a
segmenting egg, of four (or more) segments, we find in like manner, in
the great majority of cases, if not in all, that the same principle
is still exemplified; the four segments do not meet in a common
centre, but each cell is in contact with two others, and the three,
and only three, common boundary walls meet at the normal angle of
120°. A so-called _polar furrow_[358], the visible edge of a vertical
partition-wall, joins (or separates) the two triple contacts, precisely
as in Fig. 116, B.
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