But now that we have seen how a certain limited number of types of
eight-celled segmentation (or of arrangements of eight cell-partitions)
appear and reappear, here and there, throughout the whole world of
organisms, there still remains the very important question, whether
_in each particular organism_ the conditions are such as to lead to
one particular arrangement being predominant, characteristic, or even
invariable. In short, is a particular arrangement of cell-partitions to
be looked upon (as the published figures of the embryologist are apt to
suggest) as a _specific character_, or at least a constant or normal
character, of the particular organism? The answer to this question
is a direct negative, but it is only in the work of the most careful
and accurate observers that we find it revealed. Rauber (whom we have
more than once had occasion to quote) was one of those embryologists
who recorded just what he saw, without prejudice or preconception; as
Boerhaave said of Swammerdam, _quod vidit id asseruit_. Now Rauber has
put on record a considerable number of variations in the arrangement
of the first eight cells, which form a discoid surface about the
dorsal (or “animal”) pole of the frog’s egg. In a certain number of
cases these figures are identical with one another in type, identical
(that is to say) save for slight differences in magnitude, relative
proportions, or orientation. But I have selected (Fig. 168) six
diagrammatic figures, which are all _essentially different_, and these
diagrams seem to me to bear intrinsic evidence of their accuracy: the
curvatures of the partition-walls, and the angles at which they meet
agree closely with the requirements of theory, and when they depart
from theoretical symmetry they do so only to the slight extent which
we should naturally expect in a material and imperfectly homogeneous
system[395]. {381}
[Illustration: Fig. 168. Various modes of grouping of eight cells, at
the dorsal or epiblastic pole of the frog’s egg. (After Rauber.)]
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