Of these six illustrations, two are exceptional. In Fig. 168, 5, we
observe that one of the eight cells is surrounded on all sides by the
other seven. This is a perfectly natural condition, and represents,
like the rest, a phase of partial or conditional equilibrium. But it is
not included in the series we are now considering, which is restricted
to the case of eight cells extending outwards to a common boundary.
The condition shewn in Fig. 168, 6, is again peculiar, and is probably
rare; but it is included under the cases considered on p. 312, in
which the cells are not in complete fluid contact, but are separated
by little droplets of extraneous matter; it needs no further comment.
But the other four cases are beautiful diagrams of space-partitioning,
similar to those we have just been considering, but so exquisitely
clear that they need no modification, no “touching-up,” to exhibit
their mathematical regularity. It will easily be recognised that in
Fig. 168, 1 and 2, we have the arrangements corresponding to _a_
and _d_ of our diagram (Fig. 158): but the other two (i.e. 3 and 4)
represent other of the thirteen possible arrangements, which are not
included in that {382} diagram. It would be a curious and interesting
investigation to ascertain, in a large number of frogs’ eggs, all at
this stage of development, the percentage of cases in which these
various arrangements occur, with a view of correlating their frequency
with the theoretical conditions (so far as they are known, or can
be ascertained) of relative stability. One thing stands out as very
certain indeed: that the elementary diagram of the frog’s egg commonly
given in text-books of embryology,—in which the cells are depicted as
uniformly symmetrical quadrangular bodies,—is entirely inaccurate and
grossly misleading[396].
We now begin to realise the remarkable fact, which may even appear
a startling one to the biologist, that all possible groupings or
arrangements whatsoever of eight cells (where all take part in the
_surface_ of the group, none being submerged or wholly enveloped by
the rest) are referable to some one or other of _thirteen_ types or
forms. And that all the thousands and thousands of drawings which
diligent observers have made of such eight-celled structures, animal
or vegetable, anatomical, histological or embryological, are one and
all representations of some one or another of these thirteen types:—or
rather indeed of somewhat less than the whole thirteen, for there is
reason to believe that, out of the total number of possible groupings,
a certain small number are essentially unstable, and have at best, in
the concrete, but a transitory and evanescent existence.
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